Glyphidocera illiterata Meyrick, 1929

(Figs. 150–152, 281, 337, Map 27)

Glyphidocera illiterata Meyrick, 1929: 509; Becker, 1984: 41 (checklist).

Diagnosis.— Glyphidocera illiterata can be distinguished from its congeners by a unique combination of features that include a short valva, a stout digitate process of the valva, costal furcae of valva widely separate, and dorsal furca of costa about midway between digitate process of valva and ventral furca.

Description.— Head: Vertex dark brown; frontoclypeus dark brown intermixed with few pale­brown scales; outer surface of labial palpus dark brown intermixed with few pale­brown scales to near apical margin of segment II, inner margin dark brown intermixed with few pale­brown scales; scape of antenna dark brown intermixed with few palebrown scales, flagellum dark brown; male with 4th flagellum dorsally protuberant and curved, forming a deep concavity opposite slightly excavated flagellomeres 5–6; proboscis dark brown.

Thorax: Tegula and mesonotum dark brown. Legs dark brown intermixed with few pale­brown scales to near apical margins of all segments and tarsomeres. Forewing (Fig. 337): Length 5.2–7.0 mm (n = 15), dark brown intermixed with few brown and pale­brown scales; cell with one large, dark­brown spot near distal end. Undersurface dark brown. Hindwing: Pale brown basally, gradually darkening to brown towards apex.

Abdomen (Fig. 150): Male with 4 transverse, irregular rows of sex scales on intersegmental membrane between terga 2–3 and terga 3–4.

Male Genitalia (Figs. 151–152): Uncus elongate, wide at base, broadly constricted medially, broadly rounded apically, slightly keeled dorsally, apex narrowly rounded; gnathos protuberant, narrow throughout length, upturned apically; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing near midlength between posteroventral margin and base of uncus, dorsally divergent slightly and atrophied; costa of valva short, slightly sloped broadly from a shallow depression beyond a short, digitate process to dorsal furca; apex widely bifurcate, distance between furcae about as long as costa; dorsal furca curved outwardly, ventral furca twice as large as dorsal furca, and curved dorsolaterally; sacculus twisted apically slightly beyond midlength, constricting valva, widening distoventrally forming a broadly rounded cucullus; cucullus membranous and setose; vesica of aedeagus with many stout, conical cornuti, and bearing a large crescent­shaped, apical cornutus; cornutus with one side shortened near midlength, exposing a hollow, basal cavity.

Female Genitalia (Fig. 281): Papillae anales elongate, slightly wider basally than apically; apophysis posterioris and apophysis anterioris about equal in size; eighth sternum slightly narrower than eighth tergum; eighth sternum with posterior and anterior margins broadly rounded, anterior margin more emarginate medially; eighth tergum with posterior and anterior margins broadly rounded; antrum enlarged posteriorly from a constricted anterior part; posterior margin nearly straight, posterolateral margin angular; anterior part with a pair of short, spinelike processes inside cavity formed from outer wall; ductus bursae short; corpus bursae denticulate, slightly enlarged anteriorly; ductus bursae large, coiled throughout length, coils gradually decreasing in size apically; ductus seminalis with a long, 8­coiled, internal support; side opposite ductus seminalis with an elongate, platelike, internal support.

Type Examined: Holotype, ɗ, “Balboa, Panama, Canal Zone, at light, 19.6.[19]24, St. George Expedn., C.L. Collenette.” “BM Slide No. 5645.” [BMNH].

Other Specimens examined: (11 ɗ, 4 Ψ) 1 ɗ, “Est[ación] Las Pailas, 800 m, P[arque] N[acional] Rincón de la Vieja, Prov[incia] Guan[acaste], COSTA RICA, 24 Nov. a 26 Ene. 1993, J. Sihezar, G. Rodriguez, L­N­306300, 388600”, “CRI001, 211921”, “ɗ Slide No. 495”; 1 ɗ, “ 23 Oct. a 12 Nov. 1992, C. Cano”, “CRI000, 920403”, “Slide No. 500”; 1 ɗ, 1 Ψ, “Agua Buena, P. N. Guanacaste, Prov. Guana., COSTA RICA, 200 m, 7–12 Feb. 1994, E. Lopez, L­N­334800, 364100, 2692”, “CRI001, 747346”, “Ψ Slide No. 496”; “CRI001, 747377”, “Slide No. 494”; 1 ɗ, “Quepos, 80 m, P. N. Manuel Antonio, Prov. Punt., COSTA RICA, R. Zuniga, Abr. 1991, L­S­370900, 448800”, “CRI000, 593398”, “Slide No. 618”; 1 ɗ, 2 Ψ, “Fca. Cafrosa, 1300 m, Est. las Mellizas, P. N. Amistad, Prov. Punt., COSTA RICA, M. Ramirez, Mar 1991, L­S­316100, 596100”, “CRI000, 355262”, “Slide No. 682”; “CRI000, 355393”, “Ψ Slide No. 684”; “CRI000, 355278”, “Ψ Slide No. 685”; 5 ɗ, 1 Ψ, Same label data as above except, “Abr.”, “CRI000, 475034”, “Slide No. 683”; 1 ɗ, “CRI000, 474676”; 1 ɗ, “CRI000, 474630”; 1 ɗ, “CRI000, 475017”; 1 ɗ, “CRI000, 475106”; 1 ɗ, “CRI000, 474553”; 1 ɗ, Same label data as above except, “ June 1991 ”, “CRI000, 663689”, (7 in INBio, 7 in USNM).

Distribution (Map 27): In Costa Rica, Glyphidocera illiterata is known from four collecting sites from the northwestern part of the country, along the Cordillera de Guanacaste, southeast to the coast in south­central Costa Rica, east to the border near Panamá, along the Cordillera de Talamanca. G. illiterata is also known in Panamá from specimens described by Meyrick (1929).