Glyphidocera ambrosiae, new species
(Figs. 231–233, 364, Map 42)
Diagnosis.— Glyphidocera ambrosiae, G. luxuriae, and G. coturnicis are similar by sharing sex scales between abdominal terga 2–3 in male, a wide gnathos, a dorsal strut of tegumen that is widened basally, a valval cucullus with a broadly rounded apicoventral margin, and costal furcae of the valva that are divergent. However, G. ambrosiae differs from the latter species by having a longer valval cucullus and a vesica of aedeagus with an apical cornutus with a bulbus base.
Description.— Head: Scales on vertex and frontoclypeus dark brown tipped with pale brown; outer surface of labial palpus dark brown intermixed with few palebrown scales to apical margin of segment II, inner margin brown intermixed with few darkbrown and palebrown scales; scape of antenna dark brown intermixed with few palebrown scales, flagellum with alternating bands of brown and palebrown scales; male 4th flagellomere dorsally protuberant and curved, forming a shallow concavity opposite slightly excavated flagellomeres 5–6; concavity nearly closed by a few setiform scales on apex of protuberance of 4th flagellomere; proboscis brown.
Thorax: Tegula and mesonotum brown intermixed with few darkbrown scales. Legs brown intermixed with palebrown scales to apical margins of all segments and tarsomeres. Forewing (Fig. 364): Length 5.8–6.9 mm (n = 19), pale brown intermixed with brown and darkbrown scales, cell and surrounding areas paler than marginal areas; cell with 2 large, darkbrown oblique spots, 1 spot near middle, 1 spot near distal end; one short, darkbrown median streak near base. Undersurface brown intermixed with palebrown scales along costa. Hindwing: Pale brown.
Abdomen (Fig. 231): Male with 4 transverse, irregular rows of sex scales on intersegmental membrane between terga 2–3.
Male Genitalia (Figs. 232–233): Uncus elongate, wide at base, broadly constricted near 1/3, broadly rounded apically, apex narrowly rounded and setose; gnathos sparsely setose, protuberant, wide throughout length, apically upturned; tegumen slightly widened basally; dorsal strut of tegumen with arms convergent, fusing about 2/3 length from a wide base, extending dorsally, gradually widening to base of uncus; costa of valva nearly straight from a shallow depression slightly beyond a straight, basal, digitate process to a gradually upturned dorsal furca; apex widely bifurcate, each furca parallel; dorsal furca large, falcate; ventral furca extending slightly beyond outer margin of cucullus; sacculus twisted apically, constricting valva near midlength, widening distoventrally forming an elongate and broadly rounded cucullus; cucullus membranous and setose; vesica of aedeagus denticulate, bearing a large, apical cornutus; cornutus apically conical with a slightly bulbous base, one side slightly shortened, exposing a hollow, basal cavity.
Female Genitalia: Unknown.
Holotype, ɗ, “Quepos, 80 m, P[arque] N[acional] Manuel Antonio, Prov[incia] Punt[arenas], COSTA RICA, R. Zuniga, Feb. 1991, LS370900, 448800”, “INBio: COSTA RICA: CRI000, 625757” [bar code label], “INBio, Genitalia Slide by D. Adamski, Sex ɗ, No. 542” [yellow label].
Paratypes (18 ɗ): 6 ɗ, Same label data as above except, “CRI000, 346910”, “Slide No. 547”; CRI000, 347099”, “Slide No. 615”; “G. Varela & R. Zuniga, Nov. 1990 ”, “CRI000, 180343 ”, “Slide No. 616”; “Dic.”, “CRI000, 228198”; “R. Zuniga, Ene.”, “CRI000, 564315”, “Slide No. 611”; “CRI000, 366886”, “Slide No. 614”; “CRI000, 228198”, “Slide No. 544”; 1 ɗ, “Est. Los Alemendros, Prov. Guana., COSTA RICA, 300 m, 4–12 Set. 1994, E. Lopez, N334800, 369800, # 3209”, “CRI002, 019698”, “Slide No. 809”; 1 ɗ, “Monumento Nacional Guayabo, Amistad, Prov. Carta., COSTA RICA, 1100 m, Jul. 1994, G. Fonseca, LN217400, 570000, # 3126”, “CRI001, 888574”, “Slide No. 511”; 2 ɗ, “Est. Bijagual, 500 m, Res. Biol. Carara, Prov. S. José, COSTA RICA, R. Zuniga, Set. 1994, LN192250, 474760”, “CRI000, 299658”, “Slide No. 515”; “CRI000, 542103”, “Slide No. 509”; 5 ɗ, “ COSTA RICA, Heredia, Est. Biol. La Selva, 50–150 m, 10°26'N, 84°01'W, Jan. 1993, INBioOET”, “ 8 Ene. 1993, Arboleda, L001”, “CRI001, 218976”, “Slide No. 864”; June 1994 ”, “ 28 Junio 1994, Bosque secundario, L/06/107”, “CRI001, 243954”, “Slide No. 988”; “ 8 Abril 1996, L/00/207, STR 2550 m ”, “CRI002, 059844”, “Slide No. 990”; “ 29 Abril 1996, L/00/229, STR 300 m ”, “CRI002, 061889 ”, “Slide No. 883”; “ 8 Abril 1996, L/00/207. STR 2550 m ”, “CRI002, 059841”, “Slide No. 989”; 1 ɗ, “Cerro Tortuguero, P. N. Tortuguero, 0–100 m, Prov. Limón, COSTA RICA, J. Solano, Abr. 1991, LN285500, 588000”, “CRI000, 444295”, “Slide No. 608; “ Jul. 1991 ”, “CRI001, 314579”, “Slide No. 604”; “R. Delgado, Oct. 1991 ”, “CRO000, 630702”, “Slide No. 603”, (12 in INBio, 6 in USNM).
Distribution (Map 42): Glyphidocera ambrosiae is known from northwestern Costa Rica along the Cordillera de Guanacaste, east to the Cordillera Central, and from the Caribbean Coast south to the Pacific Ocean.
Etymology: The species epithet ambrosiae is derived from the Latin “ambrosia,” which means “food of the gods.”