Subfam. Zanthoxyloideae A.Juss. ex Arn. [clade C].

Genera [109]: Acmadenia, Acradenia, Acronychia, Adenandra, Adiscanthus, Agathosma, Andreadoxa, Angostura, Apocaulon, Asterolasia, Balfourodendron, Boronia, Bosistoa, Bouchardatia, Brombya, Calodendrum, Casimiroa, Choisya, Conchocarpus, Chorilaena, Coatesia, Coleonema, Comptonella, Correa, Crossosperma, Crowea, Cyanothamnus, Decagonocarpus, Decatropis, Decazyx, Desmotes, Dictamnus, Dinosperma, Diosma, Diplolaena, Drummondita, Dryades, Dutailliopsis, Dutaillyea, Empleurum, Eriostemon, Ertela, Erythrochiton, Esenbeckia, Euchaetis, Euodia, Euxylophora, Fagaropsis, Flindersia, Galipea, Geijera, Geleznowia, Halfordia, Helietta, Hortia, Ivodea, Leionema, Leptothyrsa, Lubaria, Lunasia, Maclurodendron, Macrostylis, Medicosma, Megastigma, Melicope, Metrodorea, Microcybe, Muiriantha, Myrtopsis, Naudinia, Nematolepis, Neobyrnesia, Neoschmidea, Neoraputia, Orixa, Peltostigma, Perryodendron, Pentaceras, Phebalium, Phellodendron, Philotheca, Phyllosma, Picrella, Pilocarpus, Pitavia, Pitaviaster, Plethadenia, Polyaster, Ptelea, Raputia, Raputiarana, Rauia, Raulinoa, Ravenia, Raveniopsis, Rhadinothamnus, Rutaneblina, Sarcomelicope, Sheilanthera, Sigmatanthus, Skimmia, Spiranthera, Tetractomia, Tetradium, Ticorea, Toxosiphon, Vepris, Zanthoxylum, Zieria; Number of species: ca. 1700; Characteristics: Trees or shrubs, rarely subshrubs or herbs; schizogenous oil glands present; leaves alternate or opposite or rarely whorled, simple or variously compound; flowers (3–) 4–5 (–poly)- merous, haplo- or diplostemonous; gynoecium syncarpous or fused at style, carpels 3–5 (or more), ovules 1 to several per locule; fruits dehiscent, drupaceous or baccate (rarely samaras or samaroids); endosperm lacking, scanty or copious; base chromosome number x = 18. Range: temperate and tropical regions of both hemispheres with major centers of diversity on southern continents.