Canthophorus dubius sanigarum n. ssp. (Figures 1 B, 3 C, 3 D, 4 A, 4 B, 6 D – 6 I, 7 B, 8 I, 9 A, 9 B)

Type material

Holotype: Russia: Krasnodar Terr.: Krasnaya Polyana [43°40 ʹ 32.9 ʺ N 40°12 ʹ 37.9 ʺ E], “07.. ... .. V.” [2.V.1916] (Kiritshenko), J (ZISP).

Paratypes: Russia: Krasnodar Terr.: Tuapse, 11.VI. [?] (Glusow [?], coll. Kiritshenko), 1♀ (ZISP); Tuapse, Pauk Vill., “Sev.-Kav. Kraystazra” [ North-Caucasian Regional Plant Protection Station], 8.VIII.1925 (M. Korsakova), 1J (ZISP); Ashe, 23.VI., 11.VII.1913 (Nasonov), 2♀ (ZISP); Krasnaya Polyana, “07.. ... .. V.” [2.V.1916] (Kiritshenko), 4J, 1♀ (ZISP), 1J, 1♀ (MNHN); Nebug, 16.VIII.1924 (Gittermann), 1J (ZISP).

Diagnosis

Smaller. Membrane brownish, not transparent (Figure 1 B). Sensory process of paramere apically not truncated, narrowly or widely smoothly rounded (Figure 3 C, 3 D). Conjunctival ventral lobe at apex slightly widened, subrhomboid or widely rounded; denticles on its lateral walls hardly visible or absent, dorsal wall of lobe at its apex not visible in ventral view (Figure 6 D – 6 I). Length of denticle on spicule basal plate at dorsolateral lobes less relative to length of spicule: distance from apex of denticle to apex of spicule longer than length of denticle (Figure 7 B).

Variability

In specimen from Tuapse, ventral lobe wider, its apex convoluted as tube, without lateral denticles (Figure 6 G – 6 I); perhaps this is a teratism.

Measurements

Body 6.15–8.35 in length, 3.80–5.15 in width; pronotum 3.35–4.50 in width; head 1.48–1.76 in width; ocular index 3.44–4.00; length of antennal segments (I – V): 0.34–0.41: 0.50–0.66: 0.56–0.76: 0.84–1.01: 0.94–1.09.

Etymology

The subspecies epithet is given by the name of the tribe Sanigs, Σάνηγας or Σανίγες in the ancient Greek spelling, which was inhabiting the type locality of the subspecies in the Antiquity period: “...Σάνηγας δὲ καὶ Λαζοὺς ἐποίκους εἶχον τὰ χωρία...” [Memnon of Heraclea. History of Heracleia, 37]; “Ἐνθένδε εἰς Ἀχαιοῦντα στάδιοι ἑξήκοντα, ὅσπερ ποταμὸς διορίζει Ζιλχοὺς καὶ Σανίγας” (“From here [Masaitica Riv.], it is sixty stadia to Achaeunte [Ashe or Shakhe Riv.], where the river separates the Zygii and the Sanigs ”) [Arrian. Periplus Ponti Euxini, 15].

Distribution Figure 15. Krasnodar Territory of Russia (Black Sea coast and foothills).

Canthophorus melanopterus (Herrich-Schaeffer, 1835) (Figures 1 C, 1 D, 2 A, 2 B, 3 E, 4 C –4 H, 6 J –6 M, 7 C –7 F, 8 A –8 G, 9 C, 9 D, 12 B, 13 A –13 J, 14 A – 14 I)

Cydnus melanopterus Herrich Schaeffer 1835: 55 [syntypes: Spain; lost].

Sehirus dubius var. nigra Vidal 1950: 43, n. syn.

Tritomegas melanopterus niger: Stichel 1961: 675. Canthophorus melanopterus niger: Lis 1999.

Canthophorus melanopterus: Asanova 1964: 143.

Description Lateral margin of pronotum and corium with yellowish narrow stripe; anterior angles of ventrites and connexival laterotergites with yellowish spots. Metapleural evaporatorium with long and curved peritreme. Membrane brownish or whitish.

Pygophore slightly longer than wide; its lateral walls deeply concave in middle of length (Figure 2 A); dorsal infolding long, strongly concave in middle of width; dorsal wall short (Figure 2 B).

Paramere. Hypophysis curved dorsad rather less strongly than in previous species, so that apex of hypophysis not visible in outer view (Figure 3 E).

Aedeagus (Figure 4 C). Ventral lobe of conjunctiva slightly protruding beyond apical margin of theca, slightly tapering or slightly widening at apex in ventral view; apex of lobe rounded, without lateral denticles; ventral wall of lobe rooflike, strongly elevated in apical part, with longitudinal narrow carina (Figure 6 J, 6 L); in lateral view, apical part of lobe wide, carina on its ventral wall protruding beyond apical margin of lobe as denticle (Figure 6 K, 6 M); sometimes apical margin of lobe with two small notches on either side of carina (Figure 6 L, 6 M). Apical lobes of conjunctiva long, protruding beyond dorsoventral plane of apical half of conjunctiva, basally directed dorsolaterad, in middle curved posteroventrad. Vesica with more or less long and thin membranous lateral branch originating before middle of vesica at acute angle to its longitudinal axis and lying most often on its right side; pointed apex of lateral brunch directed to apex of vesica (Figure 4 D –4 H). Dorsolateral lobes of conjunctiva long, with bases not separated from each other by impression; they directed dorsoanteriad and curved to each other by apices. Inner wall of each lobe before apex with sclerotised plate passing into spicule basal plate. Apices of spicules directed to each other and dorsad. Spicule basal plate on inner end with two triangular denticles (Figure 7 C –7 F) directed dorsomediad.

Gynatrium (Figure 8 A, 8 B). Posterior pouches located behind posterior margin of gynatrial sac, approximately as long as wide or slightly wider than long in dorsal view, with rounded posterior margin. Walls of pouches thicker and more rigid than in C. dubius; inner wall of each pouch sloped and slightly concave. Each pouch containing shaped sclerite inside (Figures 8 B, 12 B, 13 A –13 J, 14 A – 14 F, 14 H, 14 I), rarely poorly visible as very thin plate along anterior inner margin of pouch (Figure 14 G). Anterior part of gynatrial sac long and wide. Spermathecal duct with very short basal part, relatively long, porrect median part, slightly more than twice as long as distal part; sometimes proximal part elongated, attaining one third of length of shortened medial part having almost spherical shape. Tubular proximal part of spermathecal capsule gyrose and clearly longer than transverse diameter of spherical distal part (Figure 9 C, 9 D). First rami widely varying in shape and width without any geographical regularity: sharply or smoothly widening anteriorly, with rectangularly or smoothly rounded or skewed anterior ends (Figure 8 C –8 G).