Staminodeus inermis, new species

Diagnosis. Staminodeus inermis is distinguished from the other species of Staminodeus by the small size, the light reddish­brown color (but see section on variation in generic description); the short distance between the eyes, the ventrally inermous prothoracic leg in males; and the fairly short frontal spine in females.

Male. Very small, length 1.2–1.3 mm, width 0.6–0.7 mm, l/ w 1.9 –2.0, light reddishbrown. Rostrum 0.3 mm, r/p 0.9–1.0. Eyes separated by distance similar to breadth of antennal club. Frons glabrate, slightly protruded. Pronotum l/ w 0.8, slightly convex. Prosternum similar in length to mesosternum, slightly convex. Profemur (Fig. 3a) slightly sinuate, f/p 1.0–1.3, greatest width near apical 2/5, ventrally inermous; protibia (Fig. 3a) subrectate, t/f 0.8–0.9, anteroventral margin inermous, apically mucronate, promucron nearly 2/3 as long as protarsal claw; mesotibia apically mucronate, mesomucron nearly 1/4 as long as mesotarsal claw. Elytron l/ w 1.3 –1.5 (N 15). Wing/body 1.3–1.5, wing l/ w 2.9 –3.7. Median lobe (Fig. 4a) l/ w 2.6 –3.0 (N 5), laterobasal sclerites apically broadened, denticulate, median sclerite plicate, apical rami basally diverging, apically fused into triangular projection, basal 1/3 of median lobe denticulate, aedeagal apodemes shorter than median lobe.

Female. Length 1.1–1.4 mm, width 0.6–0.7 mm, l/ w 1.9 –2.0. Rostrum 0.3–0.4 mm, r/p 1.0–1.2. Eyes separated by distance slightly greater than breadth of antennal club. Frontal spine nearly 1/3 as long as diameter of eye. Pronotum l/ w 0.7 –0.8. F/p 0.9– 1.3; t/f 0.7–1.0, protibia apically mucronate, promucron nearly 1/4 as long as protarsal claw. Elytron l/ w 1.3 –1.5 (N 15). Wing/body 1.4–1.5, wing l/ w 3.1 –3.6 (N

5). Spermatheca (Fig. 5a) deflexed nearly 135, apex acute.

Type Information. Male holotype ‘‘ PANAMA, Bocas del Toro, Corriente Grande, Rio Changuinola / 9 17 30 N, 82 32 41 W, H. Wolda, Feb. 28, 1980 ’’ (CWOB) ; female allotype, same label as male holotype with different date ‘‘ Feb. 12, 1980 ’’ (CWOB); male paratypes, same labels as male holotype with different dates ‘‘ Jan. 22, 1980 ’’ (AMNH, 1), ‘‘ Jan. 23, 1980 ’’ (AMNH, 2), ‘‘ Jan. 25, 1980 ’’ (AMNH,1), ‘‘ Jan. 26, 1980 ’’ (AMNH, 1; BMNH, 1), ‘‘ Jan. 29, 1980 ’’ (BMNH, 1), ‘‘ Feb. 6, 1980 ’’ (BMNH, 1), ‘‘ Feb. 8, 1980 ’’ (BMNH, 1), ‘‘ Feb. 10, 1980 ’’ (BMNH, 1), ‘‘ Feb. 11, 1980 ’’ (CMNC, 1), ‘‘ Feb. 12, 1980 ’’ (CMNC, 1), ‘‘ Apr. 3, 1980 ’’ (CMNC, 1), ‘‘ Apr. 10, 1980 ’’ (CMNC, 1), ‘‘ Apr. 12, 1980 ’’ (CMNC, 1), ‘‘ Apr. 16, 1980 ’’ (CWOB, 1); ‘‘ Panama, Bocas del Toro, Cor. Grande, Rio Changuinola, 9 17 30 N, 82 32 41 W, leg. H. Wolda, II­06 –1980 ’’ (CWOB, 2), same labels as previous specimens with different dates ‘‘ II­14 –1980 ’’ (CWOB, 1), ‘‘ II­15 –1980 ’’ (CWOB, 1; INBC, 1), ‘‘ II­17 – 1980 ’’ (INBC, 1), ‘‘ II­26 –1980 ’’ (INBC, 2), ‘‘ III­25 –1980 ’’ (INBC, 1), ‘‘ III­ 27 – 1980 ’’ (MIUP, 2), ‘‘ IV­04 –1980 ’’ (MIUP, 1), ‘‘ IV­19 –1980 ’’ (MIUP, 1), ‘‘ IV­24 –1980 ’’ (MIUP, 1; MUCR, 3), ‘‘ IV­25 –1980 ’’ (MUCR, 1), ‘‘ V­05 – 1980 ’’ (MUCR, 1); female paratypes, same labels as male holotype with different dates ‘‘ Jan. 30, 1980 ’’ (AMNH, 1), ‘‘ Feb. 2, 1980 ’’ (AMNH, 1), ‘‘ Feb. 3, 1980 ’’ (AMNH, 1), ‘‘ Feb. 4, 1980 ’’ (AMNH, 1), ‘‘ Feb. 7, 1980 ’’ (AMNH, 1), ‘‘ Feb. 12, 1980 ’’ (BMNH, 2), ‘‘ Feb. 14, 1980 ’’ (BMNH, 1), ‘‘ Feb. 25, 1980 ’’ (BMNH, 1), ‘‘ March 7, 1980 ’’ (BMNH, 1), ‘‘ Apr. 15, 1980 ’’ (CMNC, 1), ‘‘ Apr. 27, 1980 ’’ (CMNC, 1), ‘‘ Apr. 28, 1980 ’’ (CMNC, 1); ‘‘ Panama, Bocas del Toro, Cor. Grande, Rio Changuinola, 9 17 30 N, 82 32 41 W, leg. H. Wolda, II­04 – 1980 ’’ (CMNC, 1), same labels as previous specimen with different dates ‘‘ II­06 – 1980 ’’ (CMNC, 1), ‘‘ II­07 – 1980 ’’ (CWOB, 2), ‘‘ II­14 – 1980 ’’ (CWOB, 1), ‘‘ II­15 – 1980 ’’ (CWOB, 2; INBC, 1), ‘‘ II­21 – 1980 ’’ (INBC, 3), ‘‘ II­26 – 1980 ’’ (INBC, 1), ‘‘ III­27 – 1980 ’’ (MIUP, 1), ‘‘ III­29 – 1980 ’’ (MIUP, 2), ‘‘ IV­02 – 1980 ’’ (MIUP, 1), ‘‘ IV­20 – 1980 ’’ (MIUP, 1; MUCR, 2), ‘‘ IV­24 – 1980 ’’ (MUCR, 2), ‘‘ IV­25 – 1980 ’’ (MUCR, 1).

Etymology. Named for the male plesiomorphy of lacking ventral denticulation on the prothoracic leg— inermis signifying ‘‘unarmed’’ (Nichols 1989).

Distribution. Staminodeus inermis has been collected on the Carribean slope of Panama, Provincia Bocas del Toro, Río Changuinola (Fig. 6).

Natural History. Unknown.