Ampycella fortunata sp. nov.

Figs 10, 11, 12

Material examined.

• Holotype: ♂ (CIUQ-020635), Colombia - Valle del Cauca, Buenaventura, (3.8375, -77.2436); 56 m; 12 Nov. 2022; (A. L. García, L. Delgado-Santa leg.) . • Paratypes: • 1 ♀ (CIUQ-020636), same data as the holotype • 2 ♂♂ (CIUQ-020637; CIUQ-020638) same data as the holotype .

Diagnosis.

Ampycella fortunata sp. nov. can be distinguished from the other two species of the genus by the size ratio between the spines on the free tergites: spine on free tergite II larger than the spine on free tergite III (Figs 10A, B, F, 11A, B) (vs III larger than II). Additionally, it can be distinguished from A. frizzellae by having a low ocularium with two small tubercles (vs high ocularium with two erect spines) and the entire scutal area II (vs divided into three lobes); and from A. spiniventris for having the anterolateral part of scutal area II projected forwards, anterior to the middle part (vs anterolateral parts similar to the medial portion), and having longer dorsal spines on the coxa IV.

Description.

Measurements of body and appendages. Holotype: (CIUQ-020635) DSL = 3.5, CL = 1.5, AL = 1.6, CW = 1.6, AW = 3.2, IOD = 0.4, BaCh = 0.5, FePp = 1.1; PaPp = 0.6, TiPp = 0.6, TaPp = 0.5, ClPp = 0.3, FeL I = 1.7, FeL II = 2.8, FeL III = 2.2, FeL IV = 2.4, TiL I = 1.3, TiL II = 2.2, TiL III = 1.4, TiL IV = 1.4. Paratype: (CIUQ-020636 ♀) DSL = 3.5, CL = 1.4, AL = 2.0, CW = 1.7, AW = 3.4, IOD = 0.4, BaCh = 0.5, FePp = 1.1; PaPp = 0.6, TiPp = 0.7, TaPp = 0.5; ClPp = 0.3, Fe I = 1.4, FeL II = 2.7, FeL III = 2.1, FeL IV = 2.4, TiL I = 1.2, TiL II = 2.2, TiL III = 1.3, TiL IV = 1.5. Paratypes: (CIUQ-020637; CIUQ-020638, 2 ♂♂, min-max) DSL = 3.3-3.4, CL = 1.4-1.4, AL = 1.6-1.6, CW = 1.6-1.6, AW = 3.2-3.2, IOD = 0.4-0.4, BaCh = 0.4-0.4, FePp = 1.1-1.1; PaPp = 0.5-0.6, TiPp = 0.5-0.6, TaPp = 0.5-0.6, ClPp = 0.3-0.3, FeL I = 1.3-1.4, FeL II = 2.5-2.6, FeL III = 2.1-2.1, FeL IV = 2.2-2.3, TiL I = 1.1-1.1, TiL II = 2.1-2.1, TiL III = 1.2-1.3, TiL IV = 1.3-1.4.

Male holotype (CIUQ-020635). Dorsum (Figs 10A, B, 11A, B). DS outline type alpha. Dorsal scutum widest at scutal area II; lateral borders of dorsal scutum with yellowish granules only on the middle region. Carapace mostly smooth, with a group of granules on the posterior region. Ocularium located slightly posterior to the ozopores, with a paramedian pair of small tubercles. Mesotergum well delimited, divided into four well-marked areas: I divided medially into left and right halves, with a medial large tubercle on each half and one small lateral tubercle and two anterior granules on each side; II and III entire, with a paramedian pair of large tubercles and two to three pairs on each side; IV divided into two halves, with four tubercles on each side. Posterior border of the DS substraight, with a row of tubercles. Free tergites I with a row of tubercles; II and III with a medial large spine (spine in II larger than in III) and some lateral tubercles, some of them yellowish colored.

Venter (Figs 10B, C, 11B). Stigmatic area with the posterior border convex, unarmed, only with minute granules sparsely distributed. Stigmata large, oval, and subparallel. Coxa I with two rows of large tubercles of different sizes; coxa II longer than coxae I and III, II and III with two rows of low tubercles, the anterior ones more conspicuous; coxa IV strongly backward and widened, conspicuously wider than the anterior coxae, with large tubercles in the posterior and lateral regions. Free sternites each with a row of small tubercles, the lateral larger.

Chelicerae (Fig. 11A). Chelicerae not swollen. Fixed finger with five teeth. Movable finger with ten teeth, one large tooth and distal inner surface dentate. Mesal side of the base of the fixed finger and near the base of the movable finger with setiferous tubercles of different sizes.

Pedipalps (Figs 10D, E). Trochanter unarmed. Femur slightly compressed, sub straight in lateral view, unarmed. Patella with one large dorsodistal tubercle and some granules. Tibia shorter than femur, dorsally smooth; tibia mesal II, ectal IIi. Tarsus shorter than tibia, dorsally smooth; tarsus mesal IIi, ectal IiIiii.

Legs (Figs 10A, 11C-H). Coxa I to III with one anterior row of small tubercles; IV with five longitudinal rows and with a very large dorsodistal spiniform tubercle. Trochanters I and II with two prolateral, two retrolateral, and three ventral tubercles; III with three prolateral, three retrolateral, and three ventral tubercles; IV with three prolateral, three retrolateral, and four ventral tubercles. Tubercles of trochanter IV are more conspicuous than the others. Femora I and II without conspicuous ornamentation; femora III and IV with seven rows of spines. Patellae I-IV with small tubercles. Tibiae I-IV straight and with tubercles; IV with a row of conspicuous tubercles. Claws III and IV smooth. Ratio Fe IV/DSL = 0.69. Tarsal counts 5(3)-5(3), 10(3)-10(3), 6-6, 6-6.

Penis (Fig. 12). Ventral plate (VP) sub-rectangular with medial constriction giving it a guitar-shaped appearance; distal border straight, without distal cleft. Glans + Stylus complex columnar, both parts well differentiated; stylus normally thickened, slightly curved S-shaped. MS-A1-A3 aligned in the basal part of the VP; MS-B absent; MS-C1-C3 aligned distally, similar in size and shape; MS-D1 large, similar to MS-A, distally pointed, MS-D2 absent; MS-E1-E2 very small, near MS-C cluster.

Coloration (in alcohol) (Fig. 10A-E). Carapace, chelicerae, and legs reticulated Dark yellowish brown (78) on Strong yellowish brown (74), borders of the DS and free tergites Dark yellowish brown (78). Abdominal scutum Strong yellowish brown (74), with the scutal areas Dark yellowish brown (78); Coxae dorsally reticulated on background Dark orange yellow (72). Pedipalps, reticulated. Stigmatic area with two oval areas Dark orange yellow (72) surrounded by Dark yellowish brown (78). Tarsomeri III and IV Grayish olive (110).

Female (CIUQ-020636) (Fig. 10F). Distinguished from male by carapace broader and the coda wider, genital operculum wider, spines on coxae IV present but less conspicuous than those found in males. and distal spine of femur smaller than in males.

Distribution.

Known only from the type locality.

Etymology.

fortunata is a Latin word that means happy, lucky, and blessed. It is used in reference to the meaning of the name of the type locality Buenaventura.

Remarks.

Among the known genitalia, only Licornus tama Villarreal & Kury, 2012 and Hernandarioides plana F.O. Pickard-Cambridge, 1905 have an elongated ventral plate. However, both genera have a subtle distal cleft in the distal border of the VP, which is not the typical deep cleft seen in the subfamily (e.g., Hexabunus Roewer, 1913, Hutamaia Soares & Soares, 1977, Thaumatocranaus Roewer, 1932). Ampycella fortunata sp. nov. shares the presence of an elongated ventral plate, but it is unique in not having a distal cleft. Additionally, a pair of large, apically pointing MS-D is observed in Ampycella fortunata, which is also observed in some of the genera whose genitalia is known (e.g., Hexabunus, Hernandarioides Pickard-Cambridge, 1905).