Phasgonophora granulis Delvare sp. nov.

Figs 7A-E, 8A-D, 9A, B

Type material.

Holotype ♀: Kingdom of Saudi Arabia, Al-Baha, Al Mikhwa (Shada Al-Ala Natural Reserve) [19°50'51"N, 41°18'06.12"E, Alt. 1358 m], 14.IV.2016, e.l. Acacia, leg. D. Baiocchi [KSMA]. Paratypes: 7♀, same data as holotype [KSMA]; 2♀, same data as holotype [BMNH]; 2♀, same data as holotype [USNM]; 3♀, same data as holotype [EFC]; 2♀, same data as holotype but differing as for the coordinates [19°51'39.96"N, 41°18'15.84"E, Alt. 1248 m] and collection date, 29.III.2017 [KSMA]; 2♀, Kingdom of Saudi Arabia, Asir, Muhayil, Wadi Sabian (28 km SSE of Muhayil) [18°17'54.89"N, 42°07'41.11"E, Alt. 809 m], 05.IV.2017, e.l. Acacia, leg. D. Baiocchi [KSMA].

Diagnosis.

Gaster longer than mesosoma and acuminate, with syntergum longer than mesotibia (1.15 ×) (Fig. 9A, B); gena densely and entirely punctured (Figs 7D, 8A); occiput completely strigulose (Fig. 7A); flagellum filiform, with all flagellomeres much longer than wide, F1 2.5 × as long as wide (Fig. 7C); mesosomal dorsum somewhat flattened (Fig. 8A); pronotal collum and mesonotum cristate punctured (Fig. 8A); axillae densely setose, setation masking integument posteriorly (Fig. 7E); propodeum with sharp spiracular teeth (Fig. 7E); mesepisternum with epicnemial carina forming sharp tooth mesoventrally (Fig. 8A); fore wing with dense but short setation, and pigmented track of Rs and r-m (Fig. 8D); Gt1 with weak wrinkles dorsally (Fig. 9A); Gt6 with deep punctures and very small, hardly visible, spiracle; cerci removed from base of syntergum, situated at half of its length (Fig. 9A, B).

Etymology.

The name is chosen in reference to the secondary sculpture of the areoles on the head and mesonotum, giving to them a dull, granulose appearance (see Fig. 8B).

Condition of holotype.

Specimen glued on rectangular card. Head and mesosoma partly covered with a thin layer on the bottom of areoles; second to fifth tergites with sides wide apart from each other, probably resulting from immersion in some medium.

Description of female holotype.

Body 8.4 mm. Colour. Head and mesosoma entirely black (Fig. 7A-E), metasoma brown (Fig. 9A), with syntergum darker laterally (Fig. 9B); tegula brownish (Fig. 8A); fore and mid legs dark brown, but knees, apex of tibiae and tarsi testaceous; hind leg dark brown (Fig. 8C), ventral femoral teeth and ventral side of tibia black (Fig. 8C), tarsus lighter; antenna entirely black (Fig. 7C); wings hyaline, veins dark brown (Fig. 8D).

Head (Figs 7A, B, D, 8A). Hardly wider than mesosoma; with moderately dense setation, the setae long, thin and suberect, regularly distributed according to punctures; lower face and frons strongly convex, without preorbital ridges (Fig. 7D); vertex, frons and lower face densely punctured (Fig. 7B, D), gena more coarsely punctured (Fig. 8A); both mandibles 3-teethed (Fig. 7D), lower tooth the largest and somewhat removed from the mid one; clypeus hardly protruding at free margin (Fig. 7D); edge of oral fossa thickened (Fig. 7D); tentorial pits absent (Fig. 7B); scrobal depression entirely transversely strigose, reaching median ocellus (Fig. 7B); interantennal projection (Fig. 7B) strongly compressed laterally, narrower than antennal torulus, punctulate on front surface (one row of punctures only), narrowly produced upwards, but without flange above it; vertex with short but distinct carina behind median ocellus (Fig. 7A); POL 6 × OOL (Fig. 7A); occiput entirely strigulose, except for a row of punctures behind posterior edge of eye (Fig. 7A).

Antenna (Fig. 7C). Scape linear, its apex with level of vertex; pedicel 1.2 × as long as wide, with slight basal bottle neck; anellus slightly transverse, 0.8 × as long as wide, tapering basally; funicular segments pubescent, bearing numerous, not raised multiporous plate sensilla in several intricate rows; F1 2.2 × as long as wide, shorter than F2; F2 as long as F3; F7 1.64 × as long as wide; clava 2-segmented (suture nevertheless hardly distinct), narrowly rounded apically.

Mesosoma (Figs 7E, 8A). With setae about twice as long puncture diameter, curved and suberect; setae regularly distributed on punctures, but pronotum in front of prepectus, axillae and propodeum laterally around the spiracle, densely setose, the setae adpressed there; dorsum of mesosoma somewhat flattened (Fig. 8A), with dorsal outline of pronotal collar and mesonotum straight; punctures with secondary, very fine, sculpture on their bottom (visible only at very high magnification: 800 ×) (as in Fig. 8B), thus appearing dull; pronotal collum transversely strigose (Fig. 7E); pronotal collar and mesonotum uniformly cristate punctured, the anterior wall of punctures forming crests (Fig. 8A); pronotal collar with shallow mesal depression, its sides strongly convex (Fig. 8A); pronotal carina visible laterally, forming a tooth in dorsal view; lateral panel mostly flat, with longitudinal carinulae dorsally and raised curved carina ventrally; notauli impressed (Fig. 7E); tegula with a tuft of about 10 setae anteriorly (Fig. 8A); mesoscutellum truncate anteriorly, rounded apically, its posterior margin raised and surpassing postscutellum (Fig. 7E); propodeum strongly sloping anteriorly, more strongly so posteriorly, with sharp spiracular tooth (Fig. 7E) and two irregular costulae; mesepisternum with mesodiscrimen as faint carina all over, without ventral fovea; epicnemial carina moderately raised laterally, strongly protruding mesoventrally, appearing as a sharp tooth in lateral view (Fig. 8A); ventral shelf in mesepisternum very weakly sculptured; adscrobal area, mesepimeron, and metepimeron coarsely areolate, the later bearing long setae; femoral depression of mesepisternum with only a few low carinae (Fig. 8A).

Wings (Fig. 8D). Fore wing densely setose but bare on basal cell, basal and cubital folds, marginal cell with a single, incomplete row of setae on the underside; setae generally very short on the disc of the wing, somewhat longer below MGV, PMV and Rs track; setae uniformly short on the underside of wing; MGV 0.27 × as long as SMV; PMV 0.38 × as long as MGV; STV 0.33 × as long as PMV; hind wing with 4 hamuli, the basal one the largest, removed from the followings.

Legs (Fig. 8C). Procoxa with deep depression anteriorly, margined dorsolaterally with carina raised into flange. Protibia with sharp, non-socketed apical spine. Mesotibia without dorsal pegs. Metacoxa sparsely punctured ventrally, densely so on outer surface of metafemur, with dense and fine setiferous punctures, ventral edge with irregular row of unequal teeth, outer ventral margin with a row of 8-10 teeth, basal tooth not prominent but wider than other teeth; inner basal tooth absent; apical truncation of metatibia forming a curved spine. Tarsi slender.

Metasoma (Fig. 9A, B). Petiole not visible dorsally. Gaster lanceolate, longer than mesosoma; Gt1 with weak wrinkles dorsally, setose laterally, the setae progressively longer towards the side (Fig. 9A); Gt2-5 smooth, with posterior rows of setiferous punctures, a tuft of sublateral setae longer (Fig. 9A); posterior margin of tergites hardly concave; penultimate tergite smooth anteriorly, with moderately coarse setiferous punctures posteriorly; spiracle hardly visible in sublateral position, its aperture much smaller than puncture diameter; syntergum elongate (Fig. 9A, B), 1.23 × as long as mesotibia, densely and deeply punctured, with dorsal median ridge (Fig. 9A); cerci removed from base of syntergum, situated at half of its length.

Male. Unknown.

Distribution.

Known from Saudi Arabia only in Al-Baha and Asir Regions (Fig. 17).

Host.

Anthaxia (Haplanthaxia) abdita Bílý, 1982 and A. (H.) kneuckeri ssp. zabranskyi Bílý, 1995 ( Coleoptera, Buprestidae).