Tribe Anchonieae

Matthiola incana, M. livida and M. longipetala exhibited number of primitive characters viz. continuous siphonostele, ramified sepal median bundles, sepal marginal bundles, dependent petal supply derived from one source (complex), six staminal bundles to six fertile stamens, outer staminal bundles from central stele, fused septum, lateral carpellary bundles derived from the central stele, many ovules and vascularized nectaries. This is in agreement withAl-Shehbaz (1989). For all the foregoing characters, tribe Anchonieae occupied a less advanced level (Figs 6; 7).