Virachola F. Moore, 1881 is a valid genus distinct from Deudorix Hewitson, 1863

Genomic phylogeny reveals that Deudorix Hewitson, 1863 (type species Dipsas epijarbas F. Moore, 1858) is not monophyletic and species placed in this genus belong to four different clades (Fig. 24 blue, red, green, and magenta). The first is the clade with the type species of Deudorix, thus corresponding to this genus (Fig. 24 blue). This clade is sister to several genera that include Artipe Boisduval, 1870 ( Papilio amyntor Herbst, 1804, a junior homonym, the valid name of this species is Papilio eryx Linnaeus, 1771) and Capys Hewitson, 1865 (type species Papilio alpheus Cramer, 1777). The second clade, which is not monophyletic with Deudorix, corresponds to the genus Virachola F. Moore, 1881 (type species Deudorix perse Hewitson, 1863) (Fig. 24 green), which at times is considered a valid genus (Rawlins et al. 2020). Thus, our phylogeny offers decisive evidence for treating Virachola F. Moore, 1881, stat. rest. as a genus distinct from Deudorix . However, Virachola is closely related to Artipe and could potentially be treated as its subgenus—a possibility we leave for future investigation. Furthermore, only Asian species are included in Virachola . African species form a clade that is sister to Capys, and we assign them to this genus within a newly described subgenus, detailed below. This new subgenus of Capys corresponds to the third clade of species currently in Deudorix (Fig. 24 magenta). The fourth clade is sister to all members of this group of several genera (Fig. 24 red) and it corresponds to a new genus that is described next.