Epicadus pustulosus (Mello-Leitão, 1929)

Figs 14A–D, 15E

Tobias pustulosus Mello-Leitão, 1929: 86, figs 183–184. Machado et al. 2015: 572, figs 12–24. Tobias monstrosus Simon in Mello-Leitão, 1929: 87, figs 185–186.

Epicadus pustulosus (Mello-Leitão) . Machado et al. 2017: 448, figs S7D, S 8F.

Type material: Holotype (original designation): subadult female, Fonte Boa [2°28'5.16"S, 66°8'29.49"W, Amazonas, Brazil] (MNHN 4560, examined).

Other material examined. BRAZIL: Acre: 1 male, Senador Guiomard (Reserva Extrativista de Catuaba), 10°08'43"S, 67°43'29"W, 2002, E.F. Morato (IBSP 135286) ; 1 male (IBSP 135287); 1 male (IBSP 135289); 1 male (IBSP 135290); 1 female (IBSP 135291); 1 male (IBSP 135292); all same locality and collection data . Rondônia: 1 female, Porto Velho, 8°48'10"S, 63°55'05"W, 20 January 2012, D.F. Candiani (MZSP 45458) ; 1 female, 4 July 2012, A.A. Nogueira (MZSP 51068); 1 female, 8 July 2012, R.P. Indicatti (MZSP 53085.; Bahia: 1 female, Gandu, Fazenda São Rafael (cocoa forest), 13°44'38"S, 39°29'05"W, 5 November 1968 (MNRJ 1351) ; 1 male, Una, Reserva Biológica de Una, 15°12'32"S, 39°11'45"W, 15–28 November 2000, A.D. Brescovit et al. (IBSP 46728) . Espírito Santo: 1 male, São Mateus, Reserva Floresta da Cia. Vale do Rio Doce, 19°09'05"S, 40°04'14"W, July 1997, A.J. Santos (IBSP 27015) . São Paulo: 1 female, São Paulo, Parque Modelo, 22°42'24"S, 46°48'34"W, August 1958, M. Kramer (IBSP 137989) ; 1 male, Mogi das Cruzes, Manuel Ferreira, Birituba-Açu, 23°29'22"S, 46°09'59"W, May 2001, E.K. Kashimata & R. Martins (IBSP 56568) ; 1 female, Parque Estadual da Ilha do Cardoso, 25°8'1.02"S, 47°57'41.94"W, 23–25 August 1999, G. Machado (MZSP 21592) . ECUADOR: Cotopaxi: 1 male, Rio Napo, 0°59'46"S, 78°25'18"W, January 1972, Bordon (MACN 18958).

Diagnosis. Females of E. pustulosus are very similar to those of E. caudatus in their cryptic bark-dweller habitus with rugose tegument and predominant dark-brown coloration. Both species also share squared leg segments and protuberant anterior femora suffused with many conical tubercles. However, E. pustulosus can be distinguished by the yellow blister-shaped sockets on the ventral surface of femora I and II and by the anterior projection on the epigynal plate, which covers the copulatory openings entirely. Males can be easily recognized by their equal-sized, short and spiniform opisthosomal projections, by a basal indentation on the DTA and by its curvature (pointing to the dorsum of the cymbium).

Description. See Machado et al. (2015).

Distribution. BRAZIL: São Paulo, Espiírito Santo, Bahia; ECUADOR: Cotopaxi; PANAMA: Panama City, Colón (Fig. 17A).

Variation. Most of the specimens examined by Machado et al. (2015) present a predominantly brown coloration. Here, we present two additional color variations for the females of E. pustulosus: dorsum of prosoma and opisthosoma black with front and legs yellow (Figs 14A, B) and brown with distal portion of femora I and dorsum of opisthosoma yellow (Figs 14C, D).