Phylloicus maculatus (Banks)

Figs. 64, 65

Phylloicus maculatus (Banks, 1901:369) [Type locality: Mexico, Veracruz, Presidio; MCZ; female; in Heteroplectron]. — Flint 1967:19 [female, to Phylloicus]. — Holzenthal 1988:72 [distribution].

Phylloicus maculatus is distinguished by its large size, wing pattern, and by having short, single­lobed lateral coremata, without modifications of tergite IV (Fig. 64F). The wing pattern of this species is identical to that of P. elegans and lituratus . The female genitalia are indistinguishable from those of the latter two species.

Adult. Forewing length 13.415. 4 mm, n = 31.

Head dark brown, setal warts very dark. Maxillary palps dark brown, covered with golden brown setae on basal three segments. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax dark brown, mesoscutellum golden brown; ventrolateral thorax golden. Femora golden; tibiae golden; tarsi dark brown. Metathoracic leg of male with posterior fringe of long setae. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band ivory, reaching posterior wing margin, at least 1/2 width of wing, an inverted V­shape; distal band ivory, beginning at anterior wing margin, at least 1/2 width of wing; with two basal stripes, ivory. Hind wing basal brush present in male and female; female brush short.

Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with expanded lateral flanges, lateral coremata; lateral coremata single­lobed and short, cylindrical. Tergum V without sclerotized modifications (Fig. 64F). Sternum VII with short, acute anteromesal process. Sternum VIII enclosing base of elongate sternum IX; posteromesal process notched (Fig. 64A, C). Tergum IX with paired mesal ridges extending 1/2 length of segment; posterior margin with round narrow mesal projection (Fig. 64B); lateral ridge absent; dorsal pleural setae approximately 5, ventral pleural setae approximately 8 (Fig. 64A). Preanal appendage longer than tergum X, but less than 11/2 times length, widest apically, setae filamentous, longer than appendage (Fig. 64A, B). Tergum X without basal lobes; basodorsal process short and digitate; basolateral processes long, length at least twice diameter; apex, in lateral view, rounded, in dorsal view, entire (Fig. 64A, B). Harpago rounded; peglike setae many, apical (Fig. 64A, C). Phallic endotheca with paired basolateral lobes, basolateral lobes tapered apically; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped (Fig. 64D, E).

Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch, or posterior margin entire; sternum VIII (Fig. 65C). Tergum IX without mesal ridge (Fig. 65B). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening (Fig. 65A). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes length approximately equal diameter (Fig. 65B). Sternum X with patches of short fine setae posterolaterally to anal opening (Fig. 65A). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite triangular; posterior end of spermatheca a sclerotized ovoid (Fig. 65A).

Material examined. GUATEMALA: Izabal: Cayuga, 15°32'00”N, 88°42'00"W, 1.ii.1915, Schaus — 1 female (NMNH) ; MEXICO: Veracruz: Presidio, 19°05'00”N, 96°58'00"W, 1.vi. holotype female (MCZ) ; Municipio Tilapan, Arroyo Quetzalan. Los Manantiales km 5 Rta. Orizaba­Zongolica, 18°47'28”N, 97°06'05"W, 1160 m, 17.iii.2000, Bueno, Barba, & Rojas — 1 female (UMSP) ; 13.vii.2001, Bueno & Barba — 12 males, 10 females (IBUNAM); — 3 males, 3 females (UMSP) .

Distribution. Guatemala, Mexico