Avima wayuunaiki sp. nov.

Figs 1, 2, 3, 6A, C

Type data.

COLOMBIA• ♂ holotype: La Guajira, Hatonuevo, Cerro Bañaderos, cueva [Luis Pablo Ojeda]; 11°7'51.5"N, 72°47'23.9"W [11.130972°, -72.789972°]; 978 m a.s.l.; 14 July 2015; CarBio Team 17 leg.; ICN-Ao-1976 • 7 ♂ 10 ♀: same data as holotype; ICN-Ao-1718 • 1 ♂ 1 ♀: same data as holotype; MUSENUV-Ar 2102 • 4 ♂ 2 ♀: same locality as holotype, vereda Bañaderos; cueva [Luis Pablo Ojeda]; 11°7'33.3"N, 72°47'06.9"W [11.125899°, -72.785241°]; 785 m a.s.l.; 19 May 2018; Miguel Gutiérrez leg.; MNRJ 283 • 2 ♀: same locality as previous; CBUDC-ARA 335.

Description.

Male Holotype (ICN-Ao-1976). Measurements. TL (ChL+DSL): 4.45, DSL: 2.66, CL: 1.07, CW: 1.79, AL: 1.59, AW: 2.21, BaCh: 0.55. Pedipalp: Tr: 0.60, Fe: 1.23, Pa: 0.72, Ti: 0.88, Ta: 0.70, Claw: 0.79. Leg I: Tr: 0.42, Fe: 4.10, Pa: 0.73, Ti: 3.19, Mt: 5.86, Ta: 1.31, TL: 15.61; Leg II: Tr: 0.57, Fe: 9.21, Pa: 0.98, Ti: 7.67, Mt: 12.40, Ta: 5.14, TL: 35.97. Leg III: Tr: 0.62, Fe: 5.92, Pa: 0.97, Ti: 3.65, Mt: 7.52, Ta: 1.58, TL: 20.26. Leg IV: Tr: 0.70, Fe: 8.65, Pa: 1.00, Ti: 4.85, Mt: 11.14, Ta: 1.82, TL: 28.16.

Dorsum. DS Epsilon type 2. Anterior and lateral margins of DS smooth. Ocularium low, smooth, and without median concavity (Fig. 1A, B, E). Mesotergum slightly delimited, divided into four smooth areas: area I divided into two halves; areas II-IV undivided (Fig. 2A). Posterior margin of scutum substraight and with few granules (Figs 1D, 2B). Free tergites I-III with some granules (Fig. 1B, D, E).

Venter. Coxa I with a longitudinal row of tubercles and one large bicapitate tubercle on the anteroproximal margin (Fig. 2C); coxae II-IV with some granules; coxa II longer than coxa I; coxa III longer than coxae I and II; coxa IV backward projected. Stigmata oval, small and transverse (Fig. 1C).

Chelicera. Chelicera swollen. BaCh quadrate in dorsal view, with well-marked bulla, three mesal, five anterior, and three ectal tubercles. Hand with setiferous tubercles of different sizes reaching the medial and posterior region. Fixed finger with the inner surface finely grooved. Movable finger with one trapezoid, small, sub-basal tooth, and with a dentate distal inner surface (Fig. 2D).

Pedipalps. All segments more slender than usual (Fig. 1B). Trochanter with one subapical tubercle on the ventral face. Femur with a ventroectal row of four setiferous tubercles (the two basalmost largest and the two distalmost medium-sized), and one large ventromesal setiferous tubercle in the apical portion (Fig. 2E). Patella with one large mesal setiferous tubercle. Tibia ectal III, mesal IIi. Tarsus ectal IIi, mesal IIi.

Legs. Increasing in thickness from leg I to leg IV, all smooth (Fig. 1A). Leg I filiform; coxae I-III with one dorsoanterior and one dorso-posterior tubercle; Coxa IV with some prolaterodistal tubercles; Fe III-IV darker than I-II. Fe IV length three and a half times DS length (Fig. 1A); Ta I-II each with one smooth claw; Ta III-IV with two subparallel smooth claws and without tarsal process. Tarsal counts: 9(3)-9(3)/19(3)-18(3)/7-6/7-7.

Penis. LP small (width twice the height) and apically depressed, with anterolateral acute corners (crescent-shaped) apically pointed (Fig. 3A-E). Malleus with two pairs of branched MS-A, being MS-A2 far from the base of the stylus; one pair of branched MS-B; MS-C absent; two pairs of medium-sized MS-D located in a keel between the dorsal region of the LP and the base of the stylus (Fig. 3A-C, E); MS-E2 large and branched, MS-E1 short and conical, located slightly distal to MS-E2 on the ventral side of the LP (Fig. 3C, D). Stylus elongated, mostly straight (concave at the apex), and surpassing the LP; dorsal keel present, smooth, elevated at the medial region, with a dorsoapical sharp projection; tip dorsally projected (Fig. 3E, F).

Color (in alcohol). DS and chelicerae Light Yellow (86). Pedipalps, coxae, and trochanters I-IV Pale Yellow (89). Femora to tarsi I-IV Dark Yellowish Brown (78) (Fig. 1).

Female. Paratype (ICN-Ao-1718). Similar to male, except for abdomen wider at areas II-III, chelicerae not swollen, and leg IV slender. Measurements: TL (Ch+DSL): 3.50, DSL: 2.35, CL: 0.96, CW: 1.61, AL: 1.39, AW: 1.96, BaCh: 0.42. Pedipalp: Tr: 0.41, Fe: 1.12, Pa: 0.65, Ti: 0.84, Ta: 0.79, Claw: 0.78. Leg I: Tr: 0.32, Fe: 4.27, Pa: 0.69, Ti: 3.19, Mt: 6.04, Ta: 1.29, TL: 15.8; Leg II: Tr: 0.39, Fe: 9.53, Pa: 0.98, Ti: 8.70, Mt: 12.14, Ta: 5.34, TL: 37.08. Leg III: Tr: 0.39, Fe: 9.36, Pa: 1.04, Ti: 5.77, Mt: 13.25, Ta: 1.51, TL: 31.32. Leg IV: Tr: 0.68, Fe: 8.70, Pa: 0.94, Ti: 4.83, Mt: 11.08, Ta: 1.80, TL: 28.03. Tarsal counts:?-6(3)/? -18(3)/6-6/7-6.

Etymology.

Wayuunaiki is the language spoken by the indigenous Wayuu people in northwestern Venezuela and northeastern Colombia on the Guajira Peninsula, where the species was collected. Noun in apposition.

Natural history.

Specimens of A. wayuunaiki sp. nov. were found inside a cave (Fig. 6B, C) in the northeastern slopes of Sierra Nevada de Santa Marta. This species lives sympatrically with Loxosceles guajira Cala-Riquelme, Gutiérrez-Estrada & Flórez, 2015 ( Sicariidae) and Trichomycterus spectrum DoNascimiento & Prada-Pedreros, 2020 ( Trichomycteridae).