Coprophanaeus (Coprophanaeus) dardanus (MacLeay, 1819)

Figs. 14–15

Comments. We have observed some characteristics in C. dardanus that differ from those cited by Edmonds & Zidek (2010). The pronotum of this species, as well as that of C. terrali, possesses a distinct midlongitudinal sulcus, contrary to Edmonds & Zidek’s observation that this sulcus was absent. The cap of the anterior metasternal angle is variable: although it is simple in many specimens, as stated by Edmonds & Zidek, many others have it strongly forked. In addition, the cap is often followed by a small fossa (such as cited by Edmonds & Zidek for C. degallieri and C. parvulus), and the metasternal disk has a weak midlongitudinal sulcus. These new observations are incorporated in our modification of Edmonds & Zidek’s key above.

Material examined. BRAZIL: ALAGOAS: Ibateguara, Aquidabã-II, 16.III.2008, B. K. C. Filgueiras col.— 1 specimen (CEMT); Ibateguara, Serra Grande, 9º00’01.4”S 35º51’19.1”W, 05.X.2011, B. K. C. Filgueras col.— 1 male and 2 females (CEMT); Ibateguara, Varjão, 19.IX.2007, B. Filgueiras col.— 5 specimens (CEMT); Ibateguara, Varjão, 12.X.2007, B. Filgueiras col.— 1 specimen (CEMT). AMAPÁ: Serra do Navio, Cava Urucum- Amapari, 0’53’6”N 51º52’53”W, IX.2000, R. Ribon col.— 1 specimen (CEMT). BAHIA: Igrapiuna, Pancada Grande, 13º48’8”S 39º10’3”W, 19.V.2010, R. Carvalho— 4 specimens (CEMT); Igrapiuna, Vila 5, 13 º48’8”S 39º10’3”W, 19.V.2010, R. Carvalho col.— 7 specimens (CEMT); Porto Seguro, RPPN Veracel, XI.2004, J.

Louzada col.— 2 specimens (CEMT); Valença, Cachoeira do Candengo, 0 3.III.1987, J. B. col.— 3 males (MNRJ). CEARÁ: Ubajara, Parque Nacional de Ubajara, Trilha da Samabaia, 830 m., 3º50’26”S 40º54’29”W, 15.II.2013, Vaz-de-Mello & Grossi cols.— 2 males and 1 female (CEMT); Ubajara, Parque Nacional de Ubajara, Trilha da Samabaia, 830 m., 3º50’26”S 40º54’29”W, 19.II.2013, Vaz-de-Mello & Grossi cols.— 1 male and 1 female (CEMT). MATO GROSSO: Querência, Fazenda São Luiz, II.2009, R. Andrade col.— 2 specimens (CEMT). MINAS GERAIS: Manhumirim, 30.III.1937, Zellibor-Hauff col.— 1 male (MNRJ); Mar de Espanha, XI.1987, Miro col.— 1 male (MNRJ); Parque Estadual do Rio Doce, 12.X.1992, J. Louzada col.— 2 specimens (CEMT); Rio Novo, 10.XII.2012, H. M. L. Advincola col.— 4 males and 2 females (CEMT); Rio Novo, 14.I.2013, H. M. L. Advincola col.— 1 female (CEMT); Viçosa, 18.XII.1990, Louzada & Mello cols.— 1 specimen (CEMT). PARÁ: Belém, IPEAN, XI.1984, N. Degallier col.— 1 specimen (CEMT); Benevides, VIII.1993, without collector— 1 specimen (CEMT); Novo Progresso, Fazenda Florentino, 7º8’51”S 55º23’38”W, I.2011, T. Pelissari col— 1 specimen (CEMT); Novo Progresso, Fazenda Florentino, 6º51’59”S 55º29’23”W, 12.I.2011, T. Pelissari col.— 6 specimens (CEMT); Óbidos, without date and collector— 1 female (MNRJ); Redenção, 7º46’S 51º58’W, XI.1998, P. Scheffler & T. Scheffler cols.— 6 specimens (CEMT); Redenção, Pinkaiti-Aik, 7º46’S 51º58’W, X.1999, P. Scheffler col.— 2 specimens (CEMT); São Félix do Xingu, Kayapó Pinkaiti Research Station, 31.X.1998, P. Scheffler col.— 1 specimen (CEMT). PERNAMBUCO: Iguarassu, RECD, 30.IV.2006, Arcanjo col.— 1 specimen (CEMT); Iguarassu, RECD, 25.V.2007, C. Costa col.— 1 specimen (CEMT). RIO DE JANEIRO: Angra dos Reis, Ilha Grande, IV.1992, Theotônio col.— 3 specimens (CEMT); Itatiaia, 700m, 19.X.1923, J. F. Zikán col.— 1 male (FIOC); Itatiaia, 700m, 29.XII.1924, J. F. Zikán col.— 1 male (FIOC); Itatiaia, 700m, 05.I.1925, J. F. Zikán col.— 1 female (FIOC); Parque Nacional do Itatiaia, Casa do Pesquisador, 810 m., 22–24.X.2010, Wallace Beiroz & Mario Cupello cols.— 1 female (MNRJ); Parque Nacional do Itatiaia, Casa do Pesquisador, 810 m., 22–25.II.2013, Mario Cupello col.— 1 female (MNRJ); Resende, Serrinha do Alambari, II.2011, U. Caramaschi & H. de Niemeyer cols.— 1 female (MNRJ); Restinga da Marambaia, 24.IV.1996, Marcela L. Monné & J. A. Giorgi cols.— 1 female (MNRJ); Restinga da Marambaia, 23º3’S, 43º59’W, 19.II.2010, W. Beiroz col.— 3 specimens (CEMT); Rio de Janeiro, Floresta da Tijuca, I.1933, C. A. Campos Seabra col.— 3 males and 1 female (MNRJ); Rio de Janeiro, Floresta da Tijuca, XII.1989, C. Godinho col. - (CEMT); Rio de Janeiro, Parque Lage, II.1992, F. Vaz-de-Mello col.— 17 specimens (CEMT). RORAIMA: Amajari, Ilha de Maracá, 09.V.2009, R. Andrade col.— 2 specimens (CEMT); Caracaraí, VII.1997, F. Vaz-de-Mello col.— 3 specimens (CEMT); Pacaraima, IX.1996, Ribeiro & Vazde-Mello cols.— 2 specimens (CEMT). SANTA CATARINA: Bombinhas, ARIE de Zimbros, 27ºS 48ºW, 14.X.2011, C. Oliveira col.— 2 specimens (CEMT); Brusque, Chácara Edith, 27º5’S 48º53’W, XII.2009, C. Marcon & M. Hernández cols.— 4 specimens (CEMT); Corupá, Anton Maller col.— 6 males and 7 females (MNRJ); Itapema, 27º5’13”S 49º35’54”W, 2010, M. Rocha & C. Marcon cols.— 2 specimens (CEMT); Itapema, 27º05’S 48º35’O, 17.I.2013, P. G. Silva col.— 2 males and 2 females (CEMT); Joinville, without date, Brückner col.— 1 male (MNRJ). SÃO PAULO: Peruíbe, 10.XII.1940, H. Zellibor col.— 1 male and 1 female (MNRJ); São Paulo, Camarujipe, 19.VII.2002, without collector— 1 specimen (CEMT).