Dardanus scutellatus (H. Milne Edwards, 1848)

(Fig. 4E)

Pagurus scutellatus H. Milne Edwards, 1848: 61 (Zanzibar, type locality); Forest, 1953b: 560–561; Fize & Serène, 1955: 189–195, Fig. 29, pl. 6 nos. 5–10.

Pagurus fabimanus – Estampador, 1937: 503; Yap-Chiongco, 1938: 199; Estampador, 1959: 50.

Dardanus scutellatus – Gherardi & McLaughlin, 1994: 641–642.

Material examined. 1 female SL 8.8 mm (ZRC), stn B7, Catarman, Panglao I., 9°35.9′N, 123°51.8′E, 4–30 m, 5.vi.2004; 1 male SL 6.0 mm (ZRC), stn D5, Pontod Islet, Panglao I., 9°33.6′N, 123°43.5′E, 0–3 m, 6.vi.2004; 1 male SL 9.7 mm (ZRC), stn M18, Gak-Ang Islet, off Panglao I., 9°33.0′N, 123°45.5′E, 0–1 m, 10 & 12.vi.2004; 1 male SL 9.8 mm (ZRC), stn D12, Panglao Channel, Tagbilaran, Panglao I., 9°38.5′N, 123°51.0′E, 2–4 m, 20.vi.2004; 1 ovig. female SL 4.5 mm, 1 female SL 5.2 mm, 2 males SL 6.3–6.5 mm (ZRC), stn M58, Balicasag I., 9°31.3′N, 123°41.0′E, depth unknown, 4.vii.2004; 2 ovig. females SL 5.1–7.7 mm, 5 males SL 4.7–8.9 mm (ZRC), stn M51, Mayacabac, Panglao I., 9°36.8′N, 123°52.2′E, 0 m, 30.vi.2004; 5 females SL 3.1–9.1 mm, 5 males SL 4.0– 6.8 mm (ZRC), stn R50, Looc, Panglao I., 9°35.7′N, 123°44.4′E, 3–7 m, 17.vi.2004 .

Habitat. Intertidal to shallow subtidal, 0 to approximately 5 m deep, seagrass beds and reef slopes, on sandy and soft substrates.

Remarks. This species resembles D. guttatus, but is distinctively different in colouration (see ‘remarks’ under latter species) and in habitat. It is also affiliated to D. woodmasoni, but can be distinguished by the lower margin of the big chela, which is smooth or poorly denticulated, while it is obviously denticulated in D. woodmasoni . Dardanus scutellatus is a relatively uncommon species.