Calvariopsis brasiliense (Pic, 1916) comb. nov.

(Figs 1R, 16B, 21)

= Prionocyphon brasiliensis Pic, 1916: 7

Type material. Holotype, male (MNHN): “Blumenau \ Brazil ” [handwritten]; “Type” [yellow label, handwritten]; “TYPE” [red label, printed]; “ Prionocyphon \ brasiliensis Pic ” [in Pic’s hand]; “ HOLOTYPUS \ Prionocyphon \ brasiliensis \ Pic, 1916 ” [red label, printed].

Additional material examined. Male, female (NMPC): “ SAO PAULO \ BRAS MRÁZ LGT. \ MUS. PRA- GENSE” [printed]. Male (ZMHB): “ Brasilien \ Cubatao \ 2.XI.27 \ E. Schrim S. G.”. Female (NHM): “Ilha Santo Amaro \ nr Santos, Brazil. \ G. E. Bryant \ 11.IV.1912 ”.

Redescription. Male. Body oblong oval, convex, covered with brownish suberect setae. Body uniformly dark brown, venter brown. Head very broad, 1.8× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by 1.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 1.5–2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, almost impunctate. Angle between pronotum and elytra subtly marked in dorsal outline. Elytra oblong, without traces of longitudinal ridges, widest in the middle of their length. Sides slightly curved in apical 1/2, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 1.5× diameter of a puncture.

Male genitalia. Penis (L 0.57 mm, W 0.10 mm, Fig. 21A) with subtly curved sides, parameroids short, curved, with subtriangular apices, trigonium narrow, short, pointed at apex. Tegmen (L 0.36 mm, W 0.15 mm, Fig. 21B) Ushaped, with apodemes in basal portion, apices of parameres rounded, setose. Sternite IX (L 0.24 mm, W 0.17 mm, Fig. 21C) elongate oval, with single, relatively short apodeme, apical margin rounded, covered with short, sparse setae. Tergite VIII (L 0.39 mm, W 0.37 mm, Figs 21D, E) with deep, V-shaped emargination of apical margin, and two distinct denticles in lateral portions of apical margin, apical margin not setose, apodemes as long as apical plate, diverging. Tergite IX (L 0.33 mm, W 0.34 mm, Fig. 21F) with membranous and relatively narrow apical plate, apo- demes as long as apical plate, strongly diverging.

Female. Body (Fig. 1S) dark brown, TL (n = 1) 2.60 mm. Posterior portion of each elytron with shallow, oval depression covered with wide scales, situated close to suture, slightly behind the middle (Fig. 8B). Gonocoxites (Fig. 10 J–L) triangular, gonostyli subapical, bursal sclerite (Fig. 10M, L 0.32 mm, W 0.12 mm) elongated, anterior portion with small, subconical outgrowth, central portion covered with subtle microreticulation resembling scales, posterior portion rod-like.

Measurements and ratios. Male (n = 1) TL 2.35 mm, PL 0.50 mm, PW 1.15 mm, EL 1.96 mm, EW 1.6 mm, TL/EW 1.47, PW/PL 2.30, EL/EW 1.23, EL/PL 3.92.

Variability. Sternite IX of the specimen from the collection of NMPC has a more rounded apical plate and longer apodeme.

Distribution. Known from three localities in SE Brasil.

Remarks. The species is excluded from Prionocyphon Redtenbacher as it lacks diagnostic characters of the genus. In Prionocyphon apical labial palpomere arises from side of preapical one (in Calvariopsis gen. nov. it arises from apex), antennomere 3 equal to or shorter than antennomere 2, antennomeres 2 and 3 are together shorter than antennomere 4 (in Calvariopsis gen. nov. antennomeres 2 and 3 are together longer than antennomere 4) (Ruta 2010; Watts 2010). Features of male and female genitalia of Calvariopsis gen. nov., i.e. modifications of tergite VIII and shape of bursal sclerite, differ considerably from those known in Prionocyphon .