A. tirasensis stat. nov. Haacke, 1965

Distribution. Farm Tiras, Lüderitz District, southern Namibia (Mouton & Mostert 1985; Griffin 2003) (Fig. 4).

Remarks. Haacke (1965) originally described this form as a subspecies of A. africana, and it has maintained this rank since (Mertens 1971; Branch 1981, 1988, 1998; Onderstall 1984; Branch et al. 1988; Griffin 2003), although chiefly because this poorly known taxon has not been reviewed subsequently. Under modern species concepts, the differences between the supposed subspecies of A. africana (Haacke 1965) (see above) would generally be accepted as evidence of specific distinctness, especially in light of the large disjunctions between the forms and their likely low vagility (Mouton & Mostert 1985; Jacobsen 1997). On this basis we here recognize this taxon at the rank of full species.