Tortopsis andaki sp. nov. Figs 27-32, 33-40, 41-52

Type material.

Holotype male imago (CEBUC) from Colombia, Caquetá, Curillo, río Caquetá, 220 m, 01°01 ’44” N; 75°55 ’9” W, 20.ix.2018, LG Dias, MC Zúñiga, B Toro, JP Chaux and C Molineri cols. Paratypes: 14 male and 5 female imagos (IBN). Paratypes: 10 male 5 female imagos and 1 nymph (CEBUC); 10 male and 2 female imagos (MUSENUV), same data as holotype.

Diagnosis.

Tortopsis andaki sp. nov., known from imagos of both sexes and nymph, can be distinguished from all other species of the genus by: In adults, 1) fore wing length 9.2-9.6 mm (male), 9.5-9.7 mm (female); 2) wings hyaline slightly tinted with purplish gray, veins purplish gray; 3) parastylus curved dorsally, more markedly on apical third, with a longitudinal ventral furrow (Figs 33-34); 4) penis slender, apical spine rounded and flattened (Figs 36-38); 5) parastyli receptors on sternum VIII with V-shaped sockets (Figs 39-40); 6) head shaded gray among ocelli, occiput with small gray marks (Fig. 27). The nymphs are characterized by: 1) mandibles with 7 stout small spines on inner margin of tusk see arrows in Fig. 44; 2) occiput whitish without gray markings (Fig. 31); 3) wingbuds completely whitish (Fig. 32).

Male imago. Length (mm): body, 10.8-11.0; fore wing, 9.2-9.6; hind wing, 4.0; foreleg, 4.9-5.0; cerci, 25.0-26.0. General coloration whitish shaded dorsally with light purplish gray (Figs 27). Head. Whitish shaded with light brownish gray among ocelli and on scape and pedicel, flagellum hyaline. Thorax. Pronotum hyaline shaded with purplish gray dorsally as in Fig. 27. Mesonotum whitish yellow shaded with purplish gray along medioparapsidal sutures and contiguous area, medial zone paler. Metanotum whitish yellow shaded gray dorsally. Thoracic pleura and sterna whitish. Legs whitish shaded with light purplish gray on fore tibia and fore tarsi. Wings: Membrane hyaline very lightly tinted with purplish gray, longitudinal and cross veins purplish gray. Abdomen. Translucent white, shaded slightly but extensively with purplish gray on terga, including medial area; some markings are darker (Fig. 27) and a thin medial dark line is present along most terga; terga VIII-X shaded with darker gray. Abdominal sterna whitish; sternum IX shaded very slightly with gray on median area. Genitalia (Figs 28-29, 33-38): whitish except apex of parastylus yellowish; parastylus curved dorsally more markedly on apical third (in lateral view, Figs 28 and 34-35) and with a longitudinal ventral furrow along its entire length (Fig. 33); forceps and penis translucent white, shaded very slightly gray on forceps; penis slender, spine at apex of penis rounded and flattened (Figs 36-38). Caudal filament whitish translucent.

Female imago. Length (mm): body, 10.0-10.1; fore wing, 9.5-9.7; hind wing, 3.9-4.0; cerci, 3.0. General coloration as in male but shading more strongly marked. Head black between ocelli, with small light gray marks on occiput. Wings membrane hyaline slightly tinged with whitish yellow, veins brownish. Abdomen shaded with gray on terga. Parastyli receptors on sternum VIII with sockets (Figs 39-40) with sinuous lateral margin and acute distal corner (V-shaped).

Nymph (mature female). Length: body, 17.0 mm. General coloration yellowish white shaded with gray dorsally (Fig. 30). Head. Whitish shaded with gray among ocelli, occiput completely white without marks (Fig. 31). Antennae and mouthparts whitish except apex of tusks brownish, and spines yellowish. Finger-like gill present near base of maxillae (Figs 47-48). Mandibular tusks with 7 small stout spines on inner margin, basal to subdistal tubercle (see arrows in Fig. 44); outer margin of tusk with row of 16-17 stout spines (Figs 42-44). Thorax. Pronotum, anterior ring shaded gray widely including the anterolateral pointed projections, posterior ring as in Fig. 30, both with a median pale line. Meso- and metanotum with gray and brown marks, lighter on median area; wing buds whitish, longitudinal veins hyaline, except at the basal region, with the anal and costal margins brownish. Thoracic pleura and sterna whitish. Legs whitish, with yellowish setae and apex of tarsal claws yellowish (Figs 49-52). Abdomen. Abdominal color pattern as described for the adults (Fig. 30). Gills: vestigial gills I translucent; gills II-VII well developed, formed by a pair of large whitish lamellae, the outer (dorsal) lamellae of each pair is shaded with gray on a medio longitudinal band (Fig. 30), the inner (ventral) lamellae of each pair present only a thin brownish line along trachea. Caudal filaments yellowish white.

Egg. Subcircular, white. Length, 330-350 μm; width, 280-300 μm .

Etymology.

This species is dedicated to the Andaki indigenous people, an American ethnic group that inhabited the upper Caquetá River basin.

Distribution.

Known only from the type locality.