Genus Bertia Ancey, 1887

Bertia Ancey, 1887: 53. Baker 1941: 320, 321. Zilch 1959: 275, 276. Schileyko 2003: 1362.

Ariophanta (Bertia) Thiele, 1931: 1001.

Type species.

Helix cambojiensis Reeve, 1860, by original designation.

Remarks.

Thach (2015) recognised four nominal species in Bertia: " Bertia " brookei (Adam & Reeve, 1848), B. cambojiensis (type species), B. pergrandis (Smith, 1893) and B. setzeri Thach, 2015. Nevertheless, he had overlooked key characters of the genitalia published by Godwin-Austen (1891: pl. 6, fig. 6) and Schileyko (2003: 1345, 1346, fig. 1758). The Bornean endemic species, Helix brookei Adam & Reeve, 1848, exhibits a long and cylindrical gametolytic sac, lacks an epiphallus and amatorial organ complex, and a caudal foss and a caudal horn is absent; characters that unequivocally distinguish this species from the Dyakiidae (Godwin-Austen 1891, Laidlaw 1931, Sutcharit et al. 2012). It is clear that Helix brookei Adam & Reeve, 1848 does not belong in Bertia and we recognise the following three as Bertia species.