Pandemis heparana ([Denis and Schiffermüller])

(Figs. 3, 4, 11)

Tortrix heparana Denis and Schiffermüller, 1775: 128

Pandemis heparana: [unknown author, date] (new combination).

Teras jamaicana Walker, 1863: 291 . New synonymy.

“ Archips ” jamaicana: Powell et al. 1995: 148 (new combination).

[ Archipini unplaced] jamaicana: Brown 2005: 124 .

Type material.— Tortrix heparana [Denis and Schiffermuller], 1775: Syntype, [unknown sex]: AUSTRIA: [unknown, presumed lost or destroyed]. Teras jamaicana Walker, 1863: Holotype, ♀: JAMAICA [examined; head, abdomen missing] (BMNH).

Discussion.—Unfortunately, the head and abdomen of the holotype (Fig. 3) are missing, so our conclusions are based entirely on wing pattern, coloration, and size. No species of Pandemis are described from the Neotropics (Brown 2005), so it is unlikely that Teras jamaicana represents a good species (assuming that the “ Jamaica ” label is not a mistake). We believe it is more likely that T. jamaicana represents a brief and accidental introduction to Jamaica of a described European or North American species. Of these, P. heparana is the closest match, based on comparisons to 11 female specimens in the CUIC identified by JJD (Fig. 4). Tortrix jamaicana agrees well with the wing characters described in Mutuura (1980) and Dombroskie and Sperling (2012) for P. heparana, especially in the possession of an entirely dark brown hindwing and only faint forewing banding. The vast majority of described Pandemis species are from Madagascar and East Asia. We have not compared these species to T. jamaicana, as it is unlikely that they could have made it to Jamaica. The larvae of Pandemis heparana have been recorded from an immense variety of important agricultural and ornamental plants in its native range (Brown et al. 2008), so it is not unreasonable to assume it was introduced from Europe on imported plants, especially considering it has been introduced multiple times into North America (Mutuura 1980; JJD pers. obs.).