Terebralia duboisi (Hörnes, 1855)
(Fig. 1A)
Cerithium duboisi Hörnes, 1855: 399, pl. 42, figs 4, 5.
Terebralia duboisi – Landau et al. 2013: 53, pl. 4, fig. 12 (cum syn.).
POTENTIAL SUBJECTIVE JUNIOR SYNONYMS. — (according toŠvagrovský 1971; Landau et al. 2013; and present work) Cerithium pauli Hoernes, 1875; Potamides monregalensis Sacco, 1888; Potamides Melii Sacco, 1888; Terebralia monregalensis Sacco, 1895; Clava dollfusi Hoernes, 1901; Clava holleri Hoernes, 1901; Terebralia andrzejowskii Friedberg, 1914; Terebralia praebidentata Seneš in Steininger et al. 1971, non Cerithium (Clava) bidentatum var. abbreviata Schaffer, 1912 [likely a synonym of Granulolabium plicatum (Bruguière, 1792) according to D. Kadolsky, pers. comm. 2018].
PALAEOENVIRONMENT. — Terebralia duboisi is believed to have inhabited mudflats and brackish palaeoenvironments (Landau et al. 2013; Harzhauser et al. 2014).
DISTRIBUTION. — North-eastern Atlantic. Early Miocene and Middle Miocene of France (Cossmann & Peyrot 1922; Glibert 1949). Proto-Mediterranean Sea: Burdigalian of Italy (Sacco 1895) and Turkey (İslamoğlu & Taner 2003); Middle Miocene (Serravallian) of Turkey (Landau et al. 2013); Tortonian of southern Italy (D’Amico et al. 2012). — Paratethys Sea. Early Burdigalian (Eggenburgian) of Gauderndorf (Austria) and Vélka Čausa (Slovak Republic) (Steininger et al. 1971). An Ottnangian (mid-Burdigalian) record was reported by Rupp et al. (2008). Late Burdigalian (Karpatian) of Austria cooccurring with Terebralia lignitarum (Harzhauser 2002) . Middle Miocene (Badenian) of the Central Paratethys (Harzhauser & Piller 2007). Middle Miocene (lower Sarmatian – 12.5 Ma) of Austria (Papp 1974); virtually absent from the Sarmatian sea thereafter, it reappears during the Bessarabian (late Middle Miocene – middle Sarmatian) roughly around 11 Ma in Ukraine, Romania, Moldova (Kolesnikov 1935; Simionescu & Barbu 1940). — Early Indian Ocean. Early Miocene of Iran (Landau et al. 2013).
REMARKS
Terebralia menestrieri d’Orbigny, 1844 described from the Bessarabian (Sarmatian) of Letytschiw, Oblast Chmelnyzkyj (Ukraine) differs from T. duboisi in its smaller size and delicate sculpture. Although we regard it as a possible synonym of T. duboisi, a re-examination of the Ukrainian material would be useful for assessing the possibility that it might represent a distinct offshoot of T. duboisi .
Terebralia duboisi is among the few taxa that survive the Badenian-Sarmatian Extinction Event (Harzhauser & Piller 2007).