Tribe Heterandriini + Tribe Gambusiini + Supertribe Poeciliini

[Clade 122]

Diagnosis. Members of this clade can be diagnosed by the following not uniquely derived and/or reversed features: (1) anterior margin of frontals extend anterior by between nasals [1-0]; (2) posterior supraorbital canal (2b, 3, 4a) absent or opened, forming a shallow groove [5-0]; (3) third gonapophysis forming an angle of 10-32 degrees relative to vertebral column [55-2*]; (4) first proximal radial of dorsal fin in adult males located between neural spines of 13 th and 14 th vertebrae [62-1]; and (5) hypural plate partially fused with an elongate aperture [131-1].