Parapterogramma magnum sp. nov.

urn:lsid:zoobank.org:act: E0321F2C-9CD1-41F7-9924-0B03F56E3E8B

Figs 17–19, 40G, 42

Etymology

The species name (from the Latin ‘ magnus ’, ‘large, great, big’) is in reference to the large body size compared to the other species of Parapterogramma .

Material examined

Holotype AUSTRALIA – Queensland • ♂; Rosina Creek, 14 km SE of Millaa Millaa, Palmer Highway; 720 m a.s.l.; 24 Jun.–2 Aug. 1982; S. Peck leg.; rainforest, flight intercept trap; QMBA.

Paratypes AUSTRALIA – New South Wales • 6 ♂♂, 4 ♀♀; Bruxner Park, 8 km N of Coffs Harbour; 140 m a.s.l.; 12 Jun.–26Aug. 1982; S. Peck leg.; rainforest, flight intercept trap; AMSA • 5 ♂♂, 4 ♀♀; same collection data as for preceding; DEBU • 5 ♂♂, 3 ♀♀; Ingalla State Forest, 30 km S of Macksville; 40 m a.s.l.; 12 Jun.–26 Aug. 1982; S. Peck leg.; wet sclerophyll, flight intercept trap; AMSA • 5 ♂♂; same collection data as for preceding; DEBU . – Queensland • 2 ♂♂, 1 ♀; same collection data as for holotype; QMBA • 1 ♂, 3 ♀♀; Brisbane Forest Park, Scrub Creek Road; 27°25′41″ S, 152°50′15″ E; 29 Sep.–3 Oct. 2002; C.R. Nelson leg.; Malaise trap; QMBA • 2 ♂♂; Brisbane Forest Park, Scrub Creek Road; 27°25′41″ S, 152°50′15″ E; 29 Sep. 2002; C.R. Nelson leg.; yellow pans; QMBA • 1 ♂, 3 ♀♀; Hugh Nelson Range, 19 km NW of Millaa Millaa, near the Crater; 1080 m a.s.l.; 24 Jun.–2 Aug. 1982; S. and J. Peck leg.; flight intercept trap; DEBU • 1 ♂, 2 ♀♀; Kirra State Forest, 32 km NW of Cardwell; 800 m a.s.l.; 23 Jun.–8 Aug. 1982; S. and J. Peck leg.; rainforest, flight intercept trap; QMBA • 1 ♂, 1 ♀; Kuranda State Forest, 3 km N of Kuranda; 360 m a.s.l.; 25 Jun.–3 Aug. 1982; S. Peck leg.; rainforest, flight intercept trap; DEBU • 1 ♂; Lamington National Park, Binna Burra; 900 m a.s.l.; 23 Jun.–7 Aug. 1978; S. and J. Peck leg.; Nothofagus grove, carrion; DEBU • 2 ♀♀; Mary Cairncross Park, 7 km SE of Maleny; 900 m a.s.l.; 18 Jun.–15 Aug. 1982; S. and J. Peck leg.; rainforest, flight intercept trap; QMBA • 1 ♀; Mount Finnigan Slopes, 30 km S of Cooktown; 400 m a.s.l.; 2–5 Jun. 1982; S. and J. Peck leg.; rainforest, dung trap; DEBU • 4 ♂♂, 2 ♀♀; Mount Lewis, 20 km SW of Mossman; 900 m a.s.l.; 26 Jun.–1 Aug. 1982; S. and J. Peck leg.; rainforest, flight intercept trap; DEBU • 4 ♂♂, 3 ♀♀; same collection data as for preceding; QMBA • 7 ♂♂, 10 ♀♀; Paluma, Paluma Dam Road; 850 m a.s.l.; 22 Jun.–6 Aug. 1982; S. Peck leg.; rainforest, flight intercept trap; DEBU • 7 ♂♂, 10 ♀♀; same collection data as for preceding; QMBA • 1 ♀; Sluice Creek; 17°32′ S, 145°32′ E; 1100 m a.s.l.; 14–17 Apr. 1999; G.B. and S.R. Monteith leg.; rainforest, fungus trap; QMBA • 2 ♂♂; Yabba Creek, 7 km SW of Kenilworth; 150 m a.s.l.; 18 Jun.–15 Aug. 1982; S. and J. Peck leg.; wet sclerophyll, flight intercept trap; DEBU • 1 ♂, 1 ♀; Yabba Creek, 7 km SW of Kenilworth; 15 Aug. 1982; S. and J. Peck leg.; rainforest litter; QMBA .

Description

BODY (Fig. 17). Length 1.4–1.9 mm. Head dark brown with silver-gold microtomentum on lunule, interfrontal plates, ocellar triangle, orbital plates, and occiput; gena caramel-brown; antenna brown. Frontal width 2.5× interfrontal height. Eye height 2.5× genal height. Thorax dark brown, anepimeron with a pale ventral spot. Scutum with pale grey and brown microtomentum forming indistinct dark longitudinal stripes, lateral margins broadly pale yellow microtomentose, pale M-shaped microtomentose mark at level of posterior dorsocentral bristles; scutellum dark brown with median, grey-brown microtomentose triangle. Anterior dorsocentral bristles separated by 7–8 rows of acrostichal setulae. Legs dark brown, femora apically and tibiae proximally paler, tarsi pale brown. Male mid tibia with a stout apicoventral bristle and two rows of stout ventral setae. Wing (Fig. 40G) pattern strong. CS2 0.5× CS3. R 2+3 almost straight for most of its length, distal quarter abruptly bent towards costa, causing R 2+3 to reach costa at a ~60° angle and moderately separated from costa for most of its length; costa extending very slightly beyond apex of R 4+5 (1–2 × costal width). M 1 and M 4 both extended slightly beyond dm-m; CuA+CuP long and tubular. Halter white.

MALE ABDOMEN (Fig. 18). S5 narrow, posteromedially desclerotized with long posterolateral setae. S6 short and broadened. Cercus small, flat with a very large seta and two smaller setae, mesial edges touching each other; subepandrial sclerite broadly pentagonal and medially darkened. Surstylus large, bilobed: anterior lobe low, triangular with several ventral setae, posterior lobe large, rectangular, apically setulose. Postgonite elongate with a long, triangular lateral fin and a strong, triangular posterior extension. Phallapodeme relatively short, stout with a broad, arrow-like apical part; basiphallus narrow but long and box-like with an elongate, flattened and apically setulose epiphallus. Distiphallus stout: basal part well sclerotized and tubular, with a well-developed dorsal sclerite and separate, apically bifurcate lateral sclerites; apical part with large membranous flaps and supported by basal part.

FEMALE ABDOMEN (Fig. 19). T6 semicircular, well-developed, and long-setose along posterior margin; T7 well-developed, rectangular and long-setose along posterior margin; T8 large, slightly emarginate posteromedially with long setae in posterolateral margins. Epiproct largely reduced. S6 and S7 well-developed and rectangular; S8 large, semicircular, and uniformly long-setose, posterior edge with a broad desclerotized margin. Hypoproct semicircular, with sclerotized anterolateral edges and a membranous, setulose medial part. Spermathecae stout, spherical, and smooth, stem short and sclerotized.

Remarks

Parapterogramma magnum is externally most similar to Pa. gonitellum sp. nov., from which it differs in having a broader frons (width 2.5× height), straighter R 2+3, long and tubular CuA+CuP, and the costa only extending 1–2 costal widths beyond R 4+5.