9. Dissochaeta beccariana Cogn. in A.DC. & C.DC., Monogr. Phan. 7: 559. 1891. Map 8

Neodissochaeta magnibracteata Bakh. f., Contr. Melastom.: 142. 1943. Type: Indonesia. West Kalimantan: Soengei Kenepai, 1893, J.G. Hallier 2013 (holotype: L [L0537263]!; isotypes: BO [BO1865993, BO1865994, BO1865995]!, K [K000859505]!, SING n.v.).

Neodissochaeta beccariana (Cogn.) M.P.Nayar, Kew Bull. 20: 159. 1966.

Type.

Malaysia. Sarawak: Santubong, O. Beccari PB 2190 (lectotype, designated here: FI [FI007928, image seen]!; isolectotype: K [K000859504]!).

Description.

Climbing up to 30 m in height. Branchlets terete, 3-5 mm in diameter, glabrous, sometimes with scattered bristles; nodes swollen, with a raised interpetiolar, crest-like ridge, sparsely covered with stellate hairs; internodes 8-9 cm long. Leaves: petioles terete, 1.3-1.5 cm long, glabrous except small bristles abaxiallyly; blades broadly ovate, 11-16 × 6.2-7.5 cm, subcoriaceous, base rounded, margin entire, apex acuminate, tip up to 2 cm long; nervation with 1 pair of lateral nerves and 1 pair of intramarginal nerves; adaxially glabrous, abaxially glabrous with sparsely stellate hairs on the midrib, with a basal pair of glandular patches. Inflorescences terminal, up to 22 cm long, many-flowered; main axis glabrous to sparsely stellate-puberulous; primary axes up to 18 cm long with 4 or 5 nodes, secondary axes up to 4 cm long with 1 or 2 nodes, tertiary axes ca. 1 cm with 1 node; bracts elliptic, ca. 3 × 1.2 cm, conspicuous, glabrous, white; bracteoles ovate, conspicuous, 6-8 × 6-7 mm, glabrous, white; pedicels glabrous or sparsely stellate-puberulous, 5-6 mm long in central flowers, 2-3 mm long in lateral flowers. Hypanthium tubular, 4-7 × 2-3 mm, glabrous or stellately-puberulous; calyx lobes truncate, with or without 4 small undulate tips, ca. 0.5 mm long; petal bud conical, up to 4 mm long, apex with narrow acute tip; mature petals broadly ovate, 6-7 × 5-6 mm, reflexed, purple, base clawed to slightly cordate, apex acute. Stamens 8, subequal, filaments straight, light yellow; alternipetalous stamens with 4-5 mm long filaments, anthers curved, sickle-shaped, thecae 4-5 mm long, apex acute, purple, pedoconnective ca. 1 mm long, basal crest triangular, thin, erose, irregular margin 1-2 mm long, lateral appendages paired, filiform, 3-5 mm long; oppositipetalous stamens with 5-6 mm long filaments, bent at the attachment to anthers, anthers thick, slightly curved, hook-shaped, thecae 5-7 mm long, yellow, apex obtuse, basal crest thin, ligular, 1-2 m long, lateral appendages paired, filiform, 1-2 mm long. Ovary half as long as hypanthium, apex glabrous; style glabrous, 16-18 mm long, curved at the end, slender, pale yellow; stigma minute, pinkish; extra-ovarial chambers 8, shallow to nearly undeveloped. Fruits subglobose to ovoid, 5-8 × 3-6 mm, glabrous, with 8 slight lines along the surface; calyx lobe remnants persistent. Seeds ca. 0.75 mm long.

Distribution.

Borneo.

Ecology and habitat.

Lowland to montane dipterocarp forest in open shade at 30-1400 m elevation.

Vernacular name.

Akar (Iban).

Note.

The species is easily recognised because all parts are usually glabrous, the leaf blades abaxiallyly have a pair of glandular patches at the base and the bracts and bracteoles are white. The most similar species is D. glandulosa, also with glabrous parts and glandular patches, but it has a larger, campanulate hypanthium.

Selected specimens examined.

MALAYSIA. Sabah: Beaufort, Beaufort Rail-Line, 10 m, 22 Mar 1962, G. Mikil SAN 34540 (K, L); Keningau, Pensiangan Kayu, 17 Oct 1985, Sumbing SAN 110362 (L); Ibid., Crocker Range, 1400 m, 17 Oct 1999, S.J. Davies et al. 99128 (L); Tawau, Road Kalabakan, 26 Jul 1962, Aban Gibot SAN 30555 (K, L); Ranau, Mt. Kinabalu, Dallas, 900 m, 5 Aug 1931, J. Clemens & M.S. Clemens 30342 (BM, K, L). Sarawak: O. Beccari PB 2190 (FI, K); Sibu, Rejang, Jul 1893, G.D. Haviland 3628 (BM, K); Bario, Bukit Lawi, 1500 m, 2 Aug 1985, D. Awa & B. Lee S.50537 (L); Kuching, Gunung Penrissen, 2 May 1962, Ilias Paie S.16326 (K); Ibid., Semenggoh FR., 19 Oct 1966, Banyeng ak Nyudong S.26207 (BO, K, L); Belaga, Linau Balui, Sg. Nawai, 800 m, 5 Sep 1978, B. Lee S.40000 (K, L); Kapit, Balleh, Menyiong, Sungai Sebatu, 500 m, 12 Nov 1979, Othman bin Haron et al. S.41368 (K, L); Ibid., Wong Kijang, 25 Oct 1988, Othman et al. S.56049 (L); Limbang, Sungai Ensungei, Tg. Long Amok, 11 Sep 1980, R. George et al. S.42845 (K, L); Miri, Gunung Mulu National Park, 1400 m, 9 Mar 1990, P.C. Yii & Abu Talib S.58277 (K, L). BRUNEI. Belait: Melilas, Ulu Ingei, 30 m, 5 Mar 1996, Said BRUN 17315 (K). Temburong: Ulu Belalong, 500 m, 21 Jan 1994, J. Dransfield et al. 7392 (K, L). INDONESIA. East Kalimantan: West Kutai, Long Ibut, 130 m, 16 Aug 1925, F.H. Endert 2552 (BO, L); Ibid., Mount Kemul, 1200 m, 26 Sep 1925, F.H. Endert 3572 (BO, L). West Kalimantan: Pontianak, Gunung Bentuang, 200 m, 24 Jun 1989, J.S. Burley & Tukirin 2857 (BO, L); Sungai Kenepai, 1893, J.G. Hallier 2013 (BO, K, L); Sintang, 120 m, 18 Apr 1994, A.C. Church et al. 1000 (BO, L); Kapuas Hulu, Mendalam Rivers, 170 m, 15 Mar 2000, Albertus 45 (L).