Chydaeus loeffleri (Jedlička, 1966), stat. rest.

(Figs 3, 4, 13–30)

Harpalus loeffleri Jedlička, 1966: 238

Diagnosis. In its combination of characters, including left mandible truncate at apex and elytra not fused along suture, Ch. loeffleri is most similar to Ch. irvinei, but slightly larger (length 6.8–7.8 mm, width 3.0– 3.4 mm), dorsal microsculpture developed throughout, distinct, pronotal basal angle with a tiny acute denticle protruding laterally, elytral sutural angle narrowly rounded just at tip, metacoxa with or without an additional, somewhat large medial setigerous pore, male pro- and especially mesotarsi less widened, protarsus with only tarsomeres 2–4 and mesotarsus with only tarsomeres 2 and 3 bearing adhesive vestiture, median lobe of aedeagus relatively smaller (MLL/L 0.21–0.24 versus 0.31–0.32), slightly concave along ventral margin (lateral view) and with a wider spiny patch at middle of median lobe.

Distribution (Fig. 38, ‘red circle’ and ‘red square’). Upper sources of the Tama Koshi River in the Greater Himalaya of eastern Central Nepal and southern Tibet: Rolwaling Valley (= Himalayan longitudinal Valley in eastern Central Nepal), and Rongshar valley (= uppermost portion of the Tama Koshi Valley in southern Tibet).

Remarks. The species forms two subspecies in two geographically adjacent valley systems.