Pachygnathus nasutus sp. nov.

(Figs. 16–28)

Description. Dorsum (Figs. 16, 17). Length 400–420 μm; holotrichous number of setae per segment dorsally, ad- ditional setae of small size on ventral side, soft integument slightly undulating; sclerotic crista from naso to sensillar area, microplates of setae vi inseparably linked to crista, setae exp small, barbed, medial to anterior pair of lateral eye spots.

Venter (Fig. 21). Genital valves each with 13–15 genital setae per valve; 4 anal setae per valve.

Gnathosoma (Figs. 18–20). Chelicera beak-like with a few minute teeth on both chelae, and without cheliceral setae; subcapitulum with three pairs of setae and one pair of smooth adoral setae; four sparsely barbed palpal eupathids, clearly separate from each other.

Legs (Fig 22). Solenidial formula for tarsi, tibiae, genua and femora on legs I, II, III and IV, respectively: 2- 1-0-0, 2-2-2-0, 3\4-2-2-2, 3-0-0-0, solenidion ω2 well moved anteriorly from adaxial end of lyrifissure, famulus I peg-like distally, famulus II a tiny rod abaxially.

Larva (n=1, Figs. 27, 28)

Dorsum. Length 180 μm, holotrichous, sensillae ve and sci slightly swollen, setae longest on caudal segments.

Venter. Genital area absent, 3–4 anal setae per valve.

Gnathosoma . 1 pair of palpal eupathids.

Legs. Empodia clawless, solenidial formula for tarsi, tibiae, genua and femora on legs I, II and III, respectively: 1-1-0, 2-2-2, 3-2-2, 1-0-0.

Other life stages:

Protonymph (n=3, Figs. 25, 26), 1 genital seta per valve, subcapitulum with 2 pairs of setae and one pair of adoral setae.

Deutonymph (n=4; Fig. 24), 5 genital setae per valve.

Adult male (n=5; Fig. 23), 13−15 genital setae per valve, 10 pairs of eugenitals.

Adult female (n=7; Fig. 21), 13−15 genital setae per valve, 1 pair of eugenital setae.

Type material. Holotype female, 6 paratype females, 5 paratype males, 4 paratype deutonymphs, 3 paratype protonymphs and 1 paratype larva, from soil under Maytenus cymosa (Celastraceae), vicinity of Potchefstroom, NORTH-WEST PROVINCE, 13 April 1972, J.A. van Huyssteen. Deposited at ARC-Plant Health and Protection, Pretoria, South Africa.

Differential diagnosis. This species can easily be differentiated from the neotrichous species of the genus by having a holotrichous dorsum in all stages (Fig. 27), and femora I with 3 solenidia (instead of 2 or 5–7).

Etymology. The species name refers to the prominent naso.