Hesperis quadrangula Boiss. in Ann. Sci. Nat., Bot. ser. 2, 17: 67. 1842.

Type: “Hujus specimen cum H. tristi ex Odessa sub [Aucher-Eloy] nº 149 mixtum recepi, an ex eâdem regione oriunda est?”.

Holotypus: UKRAINE: “ Odessa ”, s.d., Aucher-Eloy 149 (G-BOIS [G00332223]; iso-: BM [BM001254062], G [G00446093]) .

Notes. – The isotype at G was part of the Moricand herbarium. Furthermore, there is no duplicate of this collection at P, and it is safe to conclude that the G-BOIS unicate is the holotype.

BOISSIER (1842a) did not provide the data on the type collection between the diagnosis and full description, as he normally did for other Aucher-Eloy novelties, but for this species the data (see above) was mentioned on the last two lines of the species entry.

The status of H. quadrangula remains unresolved, and several authors (e.g., DVOŘÁK, 1973b; CZEREPANOV, 1995; MOSYAKIN & FEDORONCHUK, 1999) did not list the species as part of the flora of Ukraine. By contrast, FOURNIER (1866) listed the name in synonymy of the Turkish-Syrian endemic H. pulmonarioides . Therefore, it is safe to conclude that Aucher-Eloy made mistake in the type locality of the species.