Isatis cordigera Boiss. in Ann. Sci. Nat., Bot. ser. 2, 17: 198. 1842.

Type: “[Aucher-Eloy] E Persiâ probabiliter sed nec locus nec numerus in Schedulâ indicantur”.

Holotypus: IRAN: sine loco, s.d., Aucher-Eloy s.n. (P [P00741483]; iso-: G-BOIS [G00332594]) .

= Isatis cappadocica subsp. besseri (Trautv.) Hedge & Lamond in Rech. f., Fl. Iran. 57: 82. 1968.

Notes. – The G-BOIS sheet consists of a handwritten label by Boissier and an envelope with only two fruits taken from the unicate P00741483 that was annotated in Boissier’s handwriting as “ Isatis cordigera! Boiss. ”. Therefore, this P specimen is the holotype .

Isatis cordigera was not listed in HEDGE (1968) in the synonymy of I. cappadocica subsp. besseri .