Isatis leuconeura Boiss. & Buhse in Nouv. Mém. Soc. Imp. Naturalistes Moscou 12: 28. 1860.

Type: “Im Joluthal des Albursgebirges, [Buhse] 18 Juni 1848 (specimen unicum)”.

Holotypus: IRAN: “ Elburs ”, s.d., Buhse 1024 (G-BOIS [G00332607]; iso-: LE [LE00012917]) .

Notes. – The LE specimen is a more complete unicate than the somewhat fragmentary material in G-BOIS.

As indicated in the protologue, the LE sheet has welldeveloped oblong auricles on the sagittate-amplexicaul cauline leaves the measurements of which were given in the original description.

The G-BOIS sheet has only reduced, non-auriculate leaves at the inflorescence branches. However, there are some problems with the label information on the G-BOIS sheet. It has a pre-printed 1847 label, just as most other Persian collection of Buhse that he collected post 1847 and sent to Boissier, and the locality as “ Persia. Elburs ” [Johu Valley] and collection number 1024, compared to the LE sheet without a collection number. Finally, the G-BOIS duplicate has better developed fruits than the LE duplicate, and the latter was not annotated by Boissier. For these reasons, the G-BOIS specimen is recognized as the holotype.

HEDGE (1968: 84) indicated that Isatis leuconeura may be conspecific with I. glauca, though further studies on mature fruiting material is needed to elucidate that.