Schinia tertia (Grote)

(Figs. 1–6, 19–20, 25–26, 35, 38, 46)

Tamila tertia Grote, 1874:212 .— Grote 1875:219.— Harvey 1875:10.

Schinia tertia: Smith 1883:231 .— Smith 1891:54.— Smith 1893:279.— Dyar 1903:189.— Holland 1903:228.— Barnes and McDunnough 1917:39.— McDunnough 1938:105.— Franclemont and Todd 1983:159.— Poole 1989:897.— Poole and Gentili 1996:772.— Hardwick 1996:176.

Schinia megarena Smith, 1906:27 .— Poole 1989:895 [synonym].— Poole and Gentili 1996:772 [synonym].— Hardwick 1996:177 [valid species]. Revised synonymy.

Schinia tertia megarena; Barnes and McDunnough 1917:39.— McDunnough 1938:105.— Franclemont and Todd 1983:159.

Diagnosis. Hardwick (1996) synonymized S. albafascia with S. tertia, but gave no reason for this decision. Schinia albafascia is distinguished by the presence of a large, black quadrate discal spot in the hindwing, which is either absent or only faint and crescentshaped in tertia . The marginal band in the hindwing of albafascia is wider and more darkly colored than that of tertia . The prominent white medial band in the forewing of albafascia is wider than in tertia . In albafascia this band is approximately 30% of the length of the forewing, measured along the posterior margin, whereas in tertia it is approximately 15%. The second sternite in the male abdomen of albafascia has a large hair pencil and scent pocket; in tertia the hair pencil is absent and the scent pocket is vestigial. In the male genitalia, the uncus is noticeably longer in tertia (Fig. 25) than in albafascia (Fig. 27), and the vesica has 2 1/2 coils (Fig. 26) compared to 3 in albafascia (Fig. 28). In the female genitalia, the papillae anales are sharply curved and have a pointed apex and in tertia (Fig. 38), and are less pointed and more broadly triangulate in albafascia (Fig. 39).

Description. Abdomen (Figs. 19–20): Second sternite with hair pencil absent and a scent pocket absent (Fig. 19) or vestigial (Fig. 20). Male genitalia (Figs. 25–26): Uncus elongate, length 0.95–1.10 mm (mean = 1.025 + 0.045; n=12); valve elongate, length 2.50–2.90 mm (mean = 2.718 + 0.118; n=11); length 9.24 + 1.02 X width (n=11); saccus V­shaped. Vesica with 2 1/2 coils. Female genitalia (Figs. 35, 38): Papillae anales elongate triangular, apex curved, pointed eighth segment with minute scobinations; seventh segment with robust, elongate setae in several rows along distal margin, evenly distributed around segment, extending above distal margin of eighth segment.

Type material. Lectotype female, of Tamila tertia Grote, in BMNH, designated by Hardwick (1996). Type locality: Texas. Not examined. Lectotype male, of Schinia megarena Smith, in AMNH, designated by Todd (1982). Type locality: Utah [no specific locality]. Not examined.

Larval food plant. Unknown.

Biology. The life history of tertia is not known, but females have been observed in Kansas, Oklahoma, and eastern Colorado resting in a head down position on early blooms of Liatris punctata Hooker var. nebraskana Gaiser (CEH). The nominate variety of L. punctata overlaps much of this range, but extends into eastern Montana and North Dakota, but no tertia have been collected from this area. A third variety, mexicana Gaiser, may account for specimens from western Texas and southern New Mexico. In areas of overlap with S. albafascia in the Colorado foothills, tertia flies two to three weeks earlier and occurs in a different habitat.

Throughout much of its range, S. tertia can be the most common heliothine at lights. It regularly occurs with other species of Schinia including S. grandimedia Hardwick, S. sexplagiata Smith, S. ciliata Smith, S. gaurae (J. E. Smith), S. coercita (Grote), and S. simplex Smith in the western part of its range. In Kansas, Oklahoma, and central Texas S. tertia flies with S. nundina (Drury), S. lynx (Guenée), S. nubila (Strecker), S. ultima (Strecker), S. saturata (Grote), S. mortua (Grote), S. chrysella (Grote), S. gaurae, and S. sordida Smith.

Flight period. Collection records indicate perhaps 2 broods. There are 3 specimens in the USNM collection from Burnet Co., Texas, collected from 1–15 April and another specimen from Brazos Co., Texas, collected the end of May. There are a few specimens from July, but the principle brood begins flying in August with the majority from mid August to early September with a steady decrease into October (Fig. 45).

Distribution (Fig. 46). From Minnesota to Illinois, Iowa, eastern Missouri, Arkansas, Mississippi, Texas, west to Idaho, Utah, and south to southern California, Arizona, and New Mexico.

Material examined. All specimens are in the National Museum of Natural History, Washington, DC, unless otherwise noted. ARIZONA: APACHE CO., Baboquivari Mts., 11 Sep. 1924 (1ɗ), male genitalia slide USNM 46844, R.R. McElvare. COCONINO CO., Colorado River, Grand Canyon National Park, mi 74.4 R, old high water, 2 Sep. 2001 (2Ψ), J. Rundall (NAU); Colorado River, Grand Canyon National Park, mi 92.3 L, new high water (1Ψ), old high water (2Ψ), 3 Sep. 2001, J. Rundall (NAU); Colorado River, Grand Canyon National Park, mi 122.8 L, old high water, 4 Sep. 2001 (1ɗ), J. Rundall (NAU); Colorado River, Grand Canyon National Park, mi 166.5 L, shore, 1 Sep. 2003 (1ɗ), R.J. Delph (NAU). MOHAVE CO., Colorado River, Grand Canyon National Park, mi 180.8 R, shore (1ɗ, 3Ψ), old high water (1ɗ), new high water (1Ψ), 2 Sep. 2003, R.J. Delph (NAU). MARICOPA CO., Wickenburg, 2000’, 25 Sep. 1980 (1Ψ), 12 Oct. 1980 (1Ψ), D. & V. Hardwick (CNC). YAVAPAI CO., Granite Dells, 4 mi N of Prescott, 28 Aug. 1970 (1Ψ), 2 Sep. 1970 (1Ψ), 6 Sep. 1970 (1Ψ), L. M. Martin (CNC); Prescott, 1– 7 Sep. (1Ψ), Barnes Collection. ARKANSAS: HEMPSTEAD CO., Hope, 11 Aug. 1926 (1ɗ), 14 Aug. 1926 (1Ψ). SEBASTIAN CO., Camp Chaffee, 4 Sep. 1944, 6 Sep. 1944, L. Banks (ChAS). WASHINGTON CO., [no specific locality], 29 Aug. 1965 (1ɗ, 1Ψ), 6 Sep. 1966 (1ɗ, 1Ψ), R. L. Brown (UAF), 6 Sep. 1966 (1ɗ), R. L. Brown (EHM), 8 Sep. 1966 (1ɗ), R. L. Brown (UAF); Fayetteville, 10 Sep. 1967, J.R. Heitzman (RH), 22 Sep. 1956 (1ɗ), W. H. Whitcomb (UAF); nr. Fayetteville, 1 Sep. 1968, J.R. Heitzman (AMNH). CALI­ FORNIA: INYO CO., Death Valley National Monument, 45 mi NW Death Valley, 1700’, 15 Oct. 1966 (3ɗ), D. F. Hardwick (CNC). SAN DIEGO CO., Campo, 23 Aug. 1931 (1ɗ, 1Ψ), E. R. Tinkham (CNC) (UMSP). VENTURA CO., Cuyama Valley, Apache Canyon 0.8 mi east of Hwy. 33, 5 Sep. 1997, T.E. Dimock (TED). COLORADO: No specific locality (1ɗ, 4Ψ), Barnes Coll. ADAMS CO., Bennett, Hwy #36 near Post Office, 5643 ft., 21 Aug. 1999 (1Ψ), 24 Aug. 1999 (2Ψ) C.E. Harp (CEH). ARAPAHOE CO., S of Manilla, gas station, Hwy. I­70 at Manilla Rd., 5663 ft., 4 Aug. 2000 (1Ψ), 15 Aug. 2000 (2Ψ), 19 Aug. 1998 (1Ψ), 21 Aug. 1999 (3Ψ), 24 Aug. 1999 (1m 1Ψ), C.E. Harp (CEH). BACA CO., Picture Canyon, Comanche National Grassland, SW of Campo, 25 Aug. 2002 (2ɗ, 4Ψ), M. G. Pogue & C. E. Harp; Picture Canyon, n. of Picnic Area, SW of Campo, 4282 ft., 26 Aug. 2002 (2ɗ, 4Ψ); Comanche National Grassland, SW of Campo, Picture Canyon parking area, 4255 ft., 29 Aug. 2002 (1m 7Ψ), C.E. Harp (CEH); Springfield, S end, Hwy #385/ 287, 4409 ft., 26 Aug. 2002 (2ɗ, 11Ψ), M. G. Pogue & C. E. Harp; Springfield, S end of town, along Hwy #385/287 at truckstop lights, 4409 ft., 28 Aug. 2002 (3ɗ, 9Ψ), 28 Aug. 2003 (6ɗ, 33Ψ), 30 Aug. 2002 (10Ψ), C.E. Harp (CEH). BENT CO., Las Animas, along Hwy. #50, downtown gas station, at lights 3 am, 3817 ft., 28 Aug. 2003 (1ɗ, 6Ψ), C. Harp (CEH). ELBERT CO., downtown Kiowa, along Hwy. #86 at Pawnee St., 6412 ft., 19 Aug. 2000 (2Ψ), C.E. Harp (CEH). FREMONT CO., Penrose gas station, 1 mi N of Hwy. #50 off Hwy. 115, 5293 ft., 7 Aug. 2001 (1ɗ, 3Ψ), 11 Aug. 2001 (1ɗ, 19Ψ), 17 Aug. 2001 (3ɗ, 9Ψ), 24 Aug. 2001, (1Ψ), C.E. Harp (CEH). LINCOLN CO., Limon gas station, S of Hwy I­70 @ exit # 359, 5356 ft., 19 Aug. 1998 (2ɗ, 13Ψ), C.E. Harp (CEH). MORGAN CO., Fort Morgan, 4310 ft., 7 Aug. 1996 (1Ψ), D.J. Wright (DJW). OTERO CO., Vogel Canyon Picnic Area, 15 mi S of La Junta, 4340 ft., 18 Aug. 1997 (2Ψ), D.J. Wright (DJW); Hadley Rest Area, along Hwy. #50, ENE of La Junta, at lights 2 am, 4620 ft., 28 Aug. 2003 (1ɗ, 1Ψ), C. Harp (CEH); Rocky Ford, along Hwy #50, downtown gas station, at lights 2 am, 4161 ft., 28 Aug. 2003, C. Harp (CEH). PROWERS CO., Holly, downtown gas station along Hwy. # 50, 3427 ft., 25 Aug. 2000 (1ɗ, 1Ψ), C.E. Harp (CEH). PUEBLO CO., Colorado City, off I­25 at mkr. # 74, 5821 ft., 25 Aug. 2001 (3Ψ), C.E. Harp (CEH); 5 mi N. Pueblo, 5100 ft., 23 Aug. 1975 (1Ψ), Lafontaine & Bowen (CNC); Pueblo West, off Hwy. # 50 at McCullough Blvd., 4968 ft. 11 Aug. 2001 (3ɗ, 2Ψ), 12 Aug. 2001 (1ɗ, 4Ψ), 25 Aug. 2001 (8Ψ), C.E. Harp (CEH); Pueblo West, off Hwy. # 50 at Purcell Rd., 4968 ft., 25 Aug. 2001 (1ɗ), C.E. Harp (CEH); Pueblo N end, Hwy. I­25 @ mkr. 102, 4705 ft., 8 Aug. 1999 (1Ψ), 24 Aug. 2001 (1ɗ, 2Ψ), C.E. Harp (CEH); W of Pueblo, 26 Aug. 1958, M. May (AMNH); E of Pueblo, along Hwy #50, S of Pueblo Airport, at gas station lights, 4584 ft., 28 Aug. 2003 (3Ψ), C. Harp (CEH). WASHINGTON CO., Akron (CSU). WELD CO., Keenesburg, off Hwy. I­76, ext 39, 4955 ft., 16 Aug. 2000 (1Ψ), C.E. Harp (CEH); Nunn, 14 Aug. 1972 (UWL). IDAHO: MINI­ DOKA CO., Rupert, 10 Aug. 1957 (1Ψ) (CNC). ILLINOIS: ADAMS CO., 3 Sep. 1945, 5 Sep. 1947, Nielsen (INHS). KANE CO., [no specific locality], 26 Aug. 1940, L. Banks (ChAS). MARSHALL CO. Lacon, 23 Aug. 1938 (1Ψ), R.M. Barnes. PEORIA CO., [no specific locality], 22 Aug.1938, 23 Aug. 1938, 3 Sep. 1941, 12 Sep. 1938, 18 Sep. 1938, F. Hasbrouck (FMNH, INHS); Main Street, 21 Aug. 1938, 23 Aug. 1938, F. Hasbrouck (INHS). PUTNAM CO. 17 Sep. 1935 (1ɗ), M.O. Glenn. IOWA: BOONE CO., Ledges State Park, 21 Aug. 1959, D. Brooks (WTAMU). JOHNSON CO., Iowa City, 18 Aug. 1898, J.B. Smith Collection (AMNH). MONONA CO., [no specific locality], 24 Aug. 1960 (ISU). STORY CO., Ames, Aug. 1910 (1Ψ). WOODBURY CO., [no specific locality], 11 Sep. 1920, C.N. Ainsley (UMSP). YUBA CO., [no specific locality], 24 Aug. 1898, 25 Aug. 1898, Wickham (ISU). KANSAS: BARBER CO., Medicine Lodge, N end of town along Hwy. # 281, 1560 ft., 23 Aug. 2000 (CEH). BARTON CO., Great Bend, 15 Sep. 1988, M. Bland (FHSU); Hoisington, 30 Aug. 1994, G. Young (FHSU). BUTLER CO., rest stop 10 mi SW of El Dorado, Hwy. 35, 25 Aug. 1986, J. K. Adams (JFS). CHASE CO., 1 mi E and 1 mi N of Strong City, 1 Sep. 1990, M. Schwilling (FHSU). CLARK CO., 10 mi N of Ashland, just S of Bluff Creek, on Liatris punctata, 6 Sep. 1992, R. Huber & G. Salisbury (RH). COMMANCHE CO., Coldwater, 3 Sep. 1994, C.A. Ely (FHSU). COWLEY CO., [no specific locality], 28 Aug. 1989, G. Young (FHSU); Arkansas City, 28 Aug. 1989, G. Young (FHSU). CRAWFORD CO., 4 mi E Pittsburgh, 1–15 Sep. 1972 (1Ψ), E.L. Todd. DOUGLAS CO., [no specific locality], Snow (SMEK); Lawrence, 3 Sep. 1987 (1Ψ), J.K. Adams (JFS). ELK CO., 2 mi N of Howard, Flint Hills (T29N R10E Sec 29, NE 1/4), 30 Aug. 1981 (1m 1Ψ), D.B. & V.N. Stallings (LGC). ELLIS CO., Hays, Fort Hays State University, 12 Aug. 1988, 21 Aug. 1989, 27 Aug. 1994, 29 Aug. 1994, 5 Sep. 1994, 6 Sep. 1992, 8 Sep. 1994, C.A. Ely, 23 Aug. 1994, 25 Aug. 1994, 28 Aug. 1994, 29 Aug. 1994, 18 Sep. 1994, P. Foster, 10–14 Sep. 1994, P. Foster & C.A. Ely, 5 Sep. 1992, G.M. Wilson (FHSU); 7.5 mi N and 1 mi E of Ellis, above East Spring Creek along 130th Ave., 7 Sep. 1994, P. Foster (FHSU). ELLSWORTH CO., 4 mi N and 5 mi W of Langley, S end of Kanopolis Lake, 2 Sep. 1984, F. Davis (FHSU). FINNEY CO., Garden City, 1 Sep. 1937 (1ɗ), Franklin (LACM). FORD CO., Ford County Lake, 20 Sep. 1982, C.C. Ely (FHSU). FRANKLIN CO., [no specific locality], 2 Aug. 1944, 7 Aug. 1947, 17 Aug. 1944, 30 Aug. 1946, W.H. Howe (CNC). GRAY CO. Montezuma, 3 Sep. 1989, C.J. Ochs (FHSU). HARVEY CO., Newton (AMNH). HASKELL CO. Sublette, 2 Sep. 1989, C.J. Ochs (FHSU). JOHNSON CO., Holliday, J.R. Heitzman (RH); Prairie Village, 31 Aug. 1979, J.R. Heitzman (RH). KINGMAN CO., Kingman WMA, G. Salisbury (FHSU). KIOWA CO., 10 mi S of Greensburg, 31 Aug. 1989, G. Salisbury (FHSU); 4 mi S of Greensburg, 28 Aug. 1991, G. Salisbury (FHSU); 4– 5 mi S of Greensburg, 1991–1995, G. Salisbury (RH). LANE CO., [no specific locality], F.X. Williams (SMEK). LABETTE CO., Oswego, 27 Aug. 1964 (1ɗ), 30 Aug. 1960 (Ψ), female genitalia slide USNM 46832. LOGAN CO., 17 mi S of Monument, 15 Aug. 1975, Lafontaine & Bowen (CNC). MCPHERSON CO., 6 mi N of Canton, Maxwell Wildlife Refuge, 14 Aug. 1991, C.J. Ochs (FHSU); 7 mi S and 2 mi E of McPherson, 31 Aug. 1991, C.J. Ochs (FHSU). MITCHELL CO., 10 mi S and 1 mi W of Beloit, 25 Aug. 1990, P. Briney (FHSU). MORRIS CO., [no specific locality], 31 Aug. 1960, R. & K. Dreisbach (MSU). MORTON CO., 5 mi N and 2.5 mi W of Wilburton, Cimarron Recreation Area, 1 Sep. 1989, C.A. Ely (FHSU). RILEY CO., Manhattan, 30 Aug. 1937 (1Ψ), 1 Sep. 1937 (1Ψ), 4 Sep. 1937 (1ɗ, 1Ψ), 7 Sep. 1937 (1ɗ). RENO CO., 4.5 mi N of Plevna, 13 July 1991, D. Zongker (FHSU). RILEY CO., Manhattan, 30 Aug. – 21 Sep. (KSU); 6.5 mi S and 5 mi W of Manhattan, 3 Sep. 1994, P. Foster (FHSU). RUSSELL CO., 6 mi N and 3 mi E of Dorrance, vic. of Wilson Lake, 1 Sep. 1992, 18 Sep. 1993, P. Briney, 4 Sep. 1994, T. Beach (FHSU); 8 mi N and 8 mi E of Dorrance, vic. of Wilson Lake, 29 Sep. 1993, P. Briney (FHSU). SALINE CO., Salina, 1 Sep. 1988, C.A. Ely (FHSU). SCOTT CO., 12 mi N of Scott, Scott Wildlife Area, 26 Aug. 1994, C.A. Ely (FHSU). SEDGWICK CO., Wichita (2ɗ, 2Ψ), H.H. Walkden. SEWARD CO., 15 mi NE of Liberal, 26 Aug. 1975, Lafontaine & Bowen (CNC). SHAWNEE CO., [no specific locality], 2 Sep. 1901 (KSU). STAFFORD CO., St. John, restaurant lights, along Hwy. # 281, 1893 ft., 23 Aug. 2000 (1Ψ), C.E. Harp (CEH). STANTON CO., 4 mi W of Johnson, at Bear Creek, 19 Aug. 1990, M. Schwilling (FHSU). SUMNER CO., [no specific locality], 31 Aug. 1942, Stallings (CMP). THOMAS CO., Colby, 17 Aug. 1990, E. Weiner (FHSU). TREGO CO., 1 mi S and 3 mi E of Wakeeney, 26 Aug. 1994, 14 Sep. 1994, C.A. Ely (FHSU). MINNE­ SOTA: POLK CO., Crookston, 2 Sep. 1937, D.G. Denning (UMSP). RAMSEY CO., St. Paul, 1 Aug. 1937, A.A. Granovsky (UMSP). MISSISSIPPI: OKTIBBEHA CO., A & M College, 1 Sep. 1931 (1ɗ), R. E. Hutchins (MSU). MISSOURI: ATCHISON CO., Tarkio Prairie Nat. Area, loess prairie, 26 Aug. 1989, J.R. Heitzman (RH). BOONE CO., Columbia, 8 Sep. 1989, H. Pavulaan (AMNH). CLAY CO., Liberty (JKA); Birmingham, 29 Aug. 1968, J.R. Heitzman (RH); Kansas City, 24 Aug. 1894, F.J. Hall (AMNH). GREENE CO., Willard, Sep. 1917 (1Ψ), A.E. Brower. JACKSON CO., Independence, Adair Park, 19 Aug. 1965 (1ɗ), 29 Aug. 1965 (4ɗ, 3Ψ), J.R. Heitzman; Independence, 27 Aug. 1965 (1Ψ), 30 Aug. 1961 (2ɗ), J.R. Heitzman. NEBRASKA: BUFFALO CO., Kearney, T8N R15W Sec. 16, 16 Aug. 1998, N.E. Dankert (NED). CASS CO. Plattsmouth, 15 Sep. 1936, R.A. Leussler (AMNH). DOUGLAS CO., [no specific locality], (CNC, INHS, UMSP). LANCASTER CO., Lincoln, 29 Aug. 1937 (3ɗ), D.B. Whelan, 31 Aug. 1937 (2ɗ). PLATTE CO., Columbus, 26 Aug. 1967, E.A. Froemel (RH). SEWARD CO., 2 mi SE Seward, 18 Sep. 1972, Lafontaine & Lewis (CNC). NEW MEXICO: No specific locality (1ɗ), G. D. Hulst; CHAVES CO., Roswell, Aug. 1922 (1ɗ), Cockerell. COLFAX CO., 6 mi s of Maxwell, Hwy. I­25 @ mkr. # 420, 6040 ft., 8 Aug. 2002 (27Ψ), 12 Aug. 2002 (3ɗ, 13Ψ), C.E. Harp (CEH). DOÑA ANA CO. Las Cruces (JKA). EDDY CO., White City, 12 Sep. 1961 (1ɗ), M. & E. Roshore, A. Blanchard (AMNH); Carlsbad, 11 Sep. 1961 (1ɗ), M. & E. Roshore. GRANT CO., Silver City, R. Leuschner (RHL). GUADALUPE CO., Santa Rosa, 10 Sep. 1952, S.K & A.H. Rindge (AMNH). HIDALGO CO., Lordsburg, 22 Aug. 1960 (1ɗ), 23 Aug. 1960 (1ɗ), 27 Aug. 1960 (3ɗ), R.R. McElvare. MCKINLEY CO., Ft. Wingate, 1– 7 Aug. (1Ψ). QUAY CO., Logan, along Hwy #54, downtown gas station lights, 3791 ft., 11 Sep. 2003 (3Ψ), C. Harp (CEH); San Jon, Hwy. #469 S of Hwy. I­I­ 40, at lights, 3989 ft., 12 Sep. 2003 (1Ψ), C. Harp (CEH). SAN MIGUEL CO., Las Vegas, S.K & A.H. Rindge (AMNH). SANDOVAL CO., Jemez Springs, 16–23 Aug. (2ɗ, 3Ψ), male genitalia slide USNM 46831. OKLAHOMA: BEAVER CO. Lake Evans Chambers, 1 Sep. 1991, J.M. Nelson (ORU). BLAINE CO., Roman Nose State Park (ORU). CIMAR­ RON CO., Black Mesa State Park, s end of Lake Carl Etling, 4296 ft., 29 Aug. 2002 (8ɗ, 24Ψ), C.E. Harp (CEH). CLEVELAND CO., Norman, 2 Sep. 1950, 5 Sep. 1950, 7 Sep. 1950, Reinthal (UOKN). COMANCHE CO., Ft. Sill, Lawton, CSU study site, West Range #W­3, blacklight trap, 17–18 Aug. 2003 (1ɗ), C. Harp & M. Garhart (CEH, CSU). COT­ TON CO., 2 mi W of Walters, roadside along Hwy. #53, Arvest Bank lights, 1008 ft., 16 Aug. 2003, 19 Aug. 2003 (2ɗ, 5Ψ), C. Harp & M. Garhart (CEH); Temple, Hwy. #5, downtown carwash at lights, 1005 ft., 19 Aug. 2003 (5Ψ), C. Harp & M. Garhart (CEH). GARFIELD CO., [no specific locality], 1960 (1ɗ, 2Ψ), J.F. Reinert; Garber, s end along Hwy. # 74, 1198 ft., 25 Aug. 2000 (1Ψ), C.E. Harp (CEH). GRANT CO., Medford, gas station, along Hwy. # 11, 1067 ft., 23 Aug. 2000 (2ɗ, 3Ψ). KIOWA CO., nr. Quartz Mtns. State Park, 2 Oct. 1971, R. Boelir (SWOSU). LOVE CO., Marietta, 9 Oct. 1991, J.K. Adams (DMNS) (JFS). OKLAHOMA CO., 3 mi sw Luther, Rustic Acres Rd. Nr. Hwy. I­ 44, 980 ft., 25 Aug. 1991 (1Ψ), 9 Sep. 1991 (9Ψ), 10 Sep. 1991 (2m 3Ψ), 11 Sep. 1991 (2Ψ), 20 Sep. 1991 (2Ψ), C.E. Harp (CEH); Luther, 9 Sep. 1991, C. Harp (RH) (JFS); Bethany, 11 Sep. 1969, R. Medrud (SNU), 25 Aug. 1990, 28 Aug. 1990, 25 Sep. 1990, 25 Aug. 1991, 23 Sep. 1990 (JFS). PAYNE CO., Stillwater, 11 Sep. 1931, E. Hixon, 20 Aug. 1985, 28 Aug. 1985, 9 Sep. 1982, 15 Sep. 1982, 16 Sep. 1982, 21 Sep. 1984, 10 Oct. 1984, D.C. Arnold, Aug. 21­Oct. 4 (OKS). POTTAWATOMIE CO., 3.5 mi s McLoud, gas station, Hwy. I­40 at McLoud Rd., mkr. # 176, 1104 ft., 24 Aug. 2000 (3Ψ), C.E. Harp (CEH). ROGER MILLS CO., Cheyenne, 7 June 1937, Standish & Kaiser (OKS). ROGERS CO., Claremore, 28 Aug. 1948, C. & P. Vaurie (AMNH). TULSA CO., Bixby, 30 Aug. 1984, 15 Sep. 1987, J.T. Criswell (OKS). WOODS CO., 14 mi N and 19 mi W of Alva, 6–15 Sep. 1976, J. Farley (NED). SOUTH DAKOTA: JACKSON CO., 19 mi E of Kadoka, Hwy. I­ 90 rest stop at mkr. #168, 20 July 2001, J.K. Adams (JKA). TEXAS: No specific locality (2Ψ), (1ɗ) B Neumoegen, (2m 7Ψ), Barnes Collection, (1Ψ) Dognin Collection, (1ɗ) E.L. Graef, (1Ψ) G.D. Hulst, (1Ψ) J. Jerrmy, (1ɗ, 1Ψ) J.R. Smith, (3ɗ, 1Ψ) O. Meske. BAY­ LOR CO., Seymour (AMNH). BEXAR CO., San Antonio, 28 Sep. 1948, K.C. Emerson (OKS). BLANCO CO., Sep. (1Ψ). BOSQUE CO., Clifton (1ɗ), C.V. Riley; Laguna Park, 24 Sep. 1970, A. Blanchard (AMNH). BRAZOS CO., College Station, 15–30 May (1Ψ), 17 Sep. 1957 (1ɗ), male genitalia slide USNM 46835. BREWSTER CO., Alpine, 1926 (1ɗ), 15–21 Apr. 1928 (1ɗ), 1–7 July 1926 (1ɗ, 1Ψ), 22–31 Aug. (1Ψ), 1–7 Sep. 1926 (3ɗ), 8–14 Sep. 1926 (1ɗ), O.C. Poling, 7–13 Sep. 1963, A. Blanchard (AMNH); Alpine, Davis Mtns., 1 Sep. 1960 (3m 2Ψ), 3 Sep. 1960 (1Ψ), 12 Sep. 1958 (4ɗ), male genitalia slide USNM 46833, R.R. McElvare; The Basin, Chisos Mtns., 15 Sep. 1958 (1ɗ), R.R. McElvare. BRISCOE CO., [no specific locality] (TLA). BROWN CO., [no specific locality] (TLA). BURNET CO., Shovel Mountain, (1ɗ, 4Ψ), 1 Apr. 1902 (1Ψ), 10 Apr. 1902 (1Ψ), 15 Apr. 1902 (1Ψ). CARSON CO., White Deer, 5 Sep. 1942 (2ɗ, 2Ψ), 6 Sep. 1942 (1ɗ), 7 Sep. 1942 (4ɗ, 2Ψ), male genitalia slide USNM 46836, 8 Sep. 1942 (2ɗ, 3Ψ), 9 Sep. 1942 (1ɗ, 1Ψ). CHAMBERS CO., Black Jack Springs (1Ψ). CHILDRESS CO., 10 mi N of Childress, 31 Aug. 1996 (1Ψ), C. Bordelon (CEH) (JFS). CORYELL CO., Camp Hood, 21 Sep. 1943, (FMNH). COTTLE CO., Wildlife Management Area (AMNH); 8 mi NW Paducah, 1800 ft., 23 Sep. 1968 (1ɗ, 1Ψ), D. F. Hardwick (MSU). CULBERSON CO., [no specific locality] (AMNH). DALLAS CO., [no specific locality] (AMNH). EL PASO CO., E. Montalla Hwy., 20–24 Sep. 1995 (1Ψ), E. C. Knudson (TLA); 10 mi E of El Paso, 9 Sep. 1991, E. Knudson (JFS). HARRIS CO., [no specific locality] (CMP). HEMP­ HILL CO., Gene Howe W.M.A., 27–28 Sep. 1987 (1Ψ), E. C. Knudson (TLA); 6 mi E Canadian, 30 Aug. 1975, Lafontaine & Bowen (CNC). JEFF DAVIS CO., Ft. Davis, 19 Aug. 1984 (1Ψ), E. C. Knudson (TLA). KERR CO., Kerrville (4ɗ, 5Ψ), Aug. 1908 (1Ψ), Sep. 1908 (1ɗ), male genitalia slide USNM 46840, 7 Sep. 1958 (3ɗ, 3Ψ), R.R. McElvare, 21 Sep. 1906 (1ɗ, 1Ψ), F.C. Pratt, Oct. 1905 (1ɗ), Oct. 1908 (1ɗ); Mountain Home, 22 Sep. 1906 (2Ψ). KIMBLE CO., [no specific locality] (TLA). LLANO CO., Buchanan Dam, 27 Sep. 1986, E. C. Knudson (TLS). MITCHELL CO., [no specific locality] (TLA). MONTAGUE CO., [no specific locality] 5 Aug. 1940 (3ɗ, 1Ψ), male genitalia slide USNM 46834, 29 Aug. 1940 (4ɗ, 6Ψ), 30 Aug. 1940 (1ɗ), 1 Sep. 1940 (1ɗ), 6 Sep. 1940 (1ɗ, 5Ψ), L.H. Birdwell; Forestburg (LACM) (AMNH); Nocona, 10 Sep. 1962 (1ɗ, 1Ψ), 19 Sep. 1962 (1ɗ), R.R. McElvare. NACOGDOCHES CO., [no specific locality] (TLA). PARKER CO., Weatherford (LACM). POTTER CO., Amarillo, 24 Aug.–10 Oct. 1977, R.E. Howard (BYU); Amarillo, campus of Amarillo College, 11 Sep. 1982, S.L. Jones (AC). PRESIDIO CO., Marfa, Davis Mtns., 11 Sep. 1960 (1ɗ, 1Ψ), R.R. McElvare. RAN­ DALL CO., Canyon, 25 Aug. 1984, T. Gray, 30 Aug. 1984, G. Johnson (WTAMU). REEVES CO., Pecos, 16 Sep. 1952 (1Ψ), 22 Sep. 1952 (1Ψ), R. Leuschner. SHACKEL­ FORD CO., [no specific locality] (TLA). SUTTON CO., [no specific locality] (TLA). TARRANT CO., Benbrook, 12 Sep. 1976 (1Ψ), E.C. Knudson. TAYLOR CO., Abilene (AMNH). TERRELL CO., 10 mi E Dryden, 28 Sep. 1980 (1Ψ), E.C. Knudson; Sanderson, 28 Sep. 1980 (1ɗ), (TLA). UVALDE CO., Sabinal, Sep. 1910 (1Ψ), Oct. 1909, 1910 (2Ψ), F.C. Pratt; Concan, 2 Oct. 1994, C. Bordelon (CWB). WEBB CO., Laredo (LACM). VAL VERDE CO., Amistad, 5 Oct. 1994, J. Slotten (JFS). WICHITA CO., Wichita Falls (AMNH). UTAH: [no specific locality], 26 Aug. 1902 (1Ψ), 30 Aug. 1902 (1ɗ), O. C. Poling. CACHE CO., Logan, 15 Aug. 1937, H. F. Thornley (USU). JUAB CO., Eureka, 16 Aug. 1911 (1ɗ), 16 Aug. 1920 (3ɗ), 16 Aug. 1921 (1Ψ), 18 Aug. 1911 (1ɗ, 1Ψ), 23 Aug. 1920 (2ɗ), 26 Aug. 1921 (1ɗ), male genitalia slide USNM 46837, T. Spalding, 14 Aug. 1920, 7 Sep. 1920, T. Spalding (BYU); Mona, 14 Aug. 1929, Pack (USU). MILLARD CO., Fillmore, 20 Aug. 1941, H. F. Thornley (USU); Lynndyl, 12 Aug. 1941, H.F. Thornley (USU). SALT LAKE CO. Salt Lake City, 11–17 Aug. 1976, Knowlton & Crowe, 16– 22 Aug. 1975, Knowlton & Stephens, 25–31 Aug. 1976, Knowlton & Crowe (USU). SANPETE CO, Mayfield, 14 Aug. 1998 (2Ψ), J.M. Johnson (CEH). TOOELE CO., Stockton, 4 Aug. 1912 (USU), 19 Aug. 1903 (1ɗ, 1Ψ), T. Spalding, 24–30 Aug (1ɗ), male genitalia slide USNM 46841, 17 Sep. (2Ψ), 24 Sep. 1904 (1Ψ), T. Spalding; Dugway Proving grounds, East Dugway Dunes, 4 Aug. 1999, R. L. Johnson (BYU). UTAH CO., Salt Lake City, 20 Aug. 1914 (1Ψ), V. Fermekes (CNC); Vineyard, 14 Aug. 1912 (1Ψ), 16 Aug. 1912 (2ɗ), (CNC), 11 Aug. 1917 (1Ψ), 16 Aug. 1912 (1ɗ, 2Ψ), male genitalia slide USNM 46817, 1 Sep. 1912 (1Ψ), T. Spalding; Spanish Fork, 16 Aug. 1936 (1Ψ), T. Spalding; Dividend, 11 Aug. (1ɗ), 19 Aug. (1ɗ), T. Spalding; Provo City County Bldg., 28 Aug. 1981, C.A. Miles (BYU); Provo, Public Works, 21 Aug. 2002 (1ɗ), R.C. Mower (CEH); American Fork, 11 Aug. 1936, H.F. Thornley (USU); Spanish Fork, 1936, G.F. Knowlton (USU). WASHINGTON CO., St. George, 22 Aug. 1941, H.F. Thornley (USU). WISCONSIN: BROWN CO., [no specific locality], 1 Aug. 1957, L.W. Griewisch (UWM). COLUMBIA CO., [no specific locality], 24 Sep. 1958, P. Smith (UWM). DANE CO., Univ. Of Wisconsin arboretum, 19 Aug. 1946 (UWM). DOOR CO., [no specific locality], 26 Aug. 1961, E.R. Oatman (UWM). DOUGLAS CO., Wascott Twp., 31 Aug. 1966, J.L. Boughner (AME). MILWAUKEE CO., [no specific locality] (MPM).

Discussion. It is curious that Hardwick (1996) synonymized albafascia with tertia when there are distinct maculation and genitalic differences, whereas he treated S. megarena Smith as a valid species when there are no genitalic differences and the variation in wing maculation that occurs in megarena is found throughout the range of tertia . Historically, megarena was always treated as a subspecies of tertia (Barnes and McDunnough 1917, Franclemont and Todd 1983). Poole (1989) synonymized all subspecies and this was repeated by Poole and Gentili (1996).

The distribution of the presumed host plant of tertia, Liatris punctata, fits well with the distribution of tertia east of the Rocky Mountains, but a different host must be used west of the Rockies. More research is needed to discover the western host plant of tertia .

The figures in Hardwick (1996) referring to S. tertia (F10, W1, W2) are actually S. albafascia .