Makromastax mandibularis (Chemsak & Linsley, 1963), comb. nov.
(Figs. 23–31)
Haplidus mandibularis Chemsak and Linsley, 1963: 229; Monné, 2017a: 239 (cat.).
Haplidus nitidus Chemsak and Linsley, 1963: 228; Monné, 2017a: 239 (cat.). S yn. nov.
Chemsak and Linsley (1963) described Haplidus nitidus and H. mandibularis in the same work, and both species from Mexico, respectively: Morelos and Puebla. Still according to them, H. mandibularis differs from H. nitidus by the prominent, bifid mandibles, narrower pronotum, and shorter antennal scape. However, the mandibles in H. mandibularis are very similar, although distinct longer in male, and the scape is also very similar in shape and length in both species. The difference in the proportions between scape and antennomere III reported in these species, is a common sexual dimorphism (scape longer than third segment [ H. nitidus]; scape subequal in length to third segment [ H. mandibularis]). The same regarding the prothoracic shape.
Based in the original descriptions, examination of photographs of the holotypes of both species, and detailed exam of a male, we concluded that H. mandibularis is the male of H. nitidus . As H. mandibularis and H. nitidus were described in the same work (Chemsak and Linsley 1963) and the Principle of Priority does not apply (ICZN 1999: Article 24.2.2), we chose Haplidus mandibularis as the valid name because is in the males that the mandibles are long and falciform, the more conspicuous characteristic of the genus. After the original descriptions, these species were only mentioned in checklists and catalogues.
Material examined. MEXICO, Guerrero: Xalitla (600 m), 1 male, 1.VIII.2011, D. Curoe col. (MZSP). New state record.