KEY TO THE SPECIES OF THE GENUS TOKTOKKUS 1. Apex of elytra not sloped (dorsal view) and depressed with clear margin (Fig. 3B)............... T. congolensis- Apex of elytra sloped (dorsal view) and flat to slightly convex (e.g. Fig. 3A) ...............................................2 2. Margin of prosternal collar expanded and folded out into a large lip (Fig. 3C); elytra round, almost heart- shaped (Fig. 3C); declivous portion of elytra with little to no tubercles..................................... T. tschinkeli- Margin of prosternal collar not expanded, only occasionally folded out; elytra round or elongate; declivous portion of elytra with tuberculate rows .........................................................................................................3 3. Disc of pronotum with prominent punctures (Fig. 3D); gold setae present on elytra (Fig. 3D)..... T. herero- Disc of pronotum only with micropunctures; elytra not covered with setae................................................44. Tubercles on elytral sides round/globular, vertically direct (Fig. 2B, C) ......................................................5- Tubercles on elytral sides pointed, directed posteriorly (e.g. Fig. 3F) ..........................................................6 5. Elytral tubercles dense (1.0–1.5 diameters apart), laterally reaching humerus; elytra round (Fig. 2B) ........................................................................................................................................................... T. mulleri - Elytral tubercles sparse (2–3 diameters apart), laterally terminating prior to humerus; elytra elongate to round (Fig. 2C)................................................................................................................................ T. schultzei6. Elytral tubercles sparse (4–6 lateral rows), deeply angled...........................................................................7- Elytral tubercles dense (more than 6 lateral rows), slightly angled (Fig. 2B–F).........................................8 7. Elytral tubercles distinct, not confluent, nearly reaching humeri (Fig. 3A) ................................ T. barclayi - Elytral tubercles small and short, confluent into rows, terminating well before humeri (Fig. 2A) ...................................................................................................................................................... T. mashunus8. Microtubercles present between tuberculate rows (e.g. Fig. 3E), tuberculate rows rarely elevated on ridges ...............................................................................................................................................................9- Elytral tubercles all relatively the same size, no microtubercles present between rows, tuberculate rows often elevated on ridges ................................................................................................................................10 9. Body size fairly small (18.0–23.0 mm); elytral tubercles relatively dense, almost confluent; tubercles relatively short .................................................................................................................... T. tuberculipennis - Body size medium to large (28.0–32.0 mm); elytral tubercles less dense, only occasionally confluent; tubercles taller ................................................................................................................................ T. makuya 10. Elytral tubercles large, often confluent (Fig. 3F); tuberculate rows extend over humeri to scutellum; humeri prominent; disk of elytra where smooth often flat (Fig. 3F) ......................................... T. waclawae - Tubercles medium sized (Fig. 2E, F), not confluent; tuberculate rows end at humeri; humeri not prominent; disc of elytra where smooth generally convex .................................................................................... T. vialis