Pteropus mariannus Desmarest, 1822 . Mammalogie, in Encycl. Méth., 2(Suppl.):547.

TYPE LOCALITY: Marianna Isis, Guam (USA) .

DISTRIBUTION: Mariana Isis and Caroline Isis (including Palau Isis), Ryukyu Isis (Japan).

STATUS: CITES - Appendix I; U.S. ESA - Endangered in Guam as P. m. mariannus; IUCN - Endangered.

SYNONYMS: keraudren, loochooensis, paganensis, pelewensis, ualanus, ulthiensis, vanikorensis, yapensis .

COMMENTS: Includes loochooensis, pelewensis, ualanus, and yapensis; see Kuroda (1938). Corbet and Hill (1980:37-38) listed these forms as distinct species without comment; but see Yoshiyuki (1989), who treated loochooensis as a separate species. Probably includes vanikorensis, see Troughton (1930).