Archaeocercoides puchkovi Simutnik sp. nov.

Figs 1A, B, D, 2, 3

Material.

Holotype, SIZK, no. UA-28099, 1♀, Varash District, Rovno Region, Ukraine; Rovno amber; late Eocene. The inclusion is in a yellow and clear piece of amber in a shape of parallelepiped (ca. 40 × 10 × 9 mm), one side of which contains a layer of organic residues. All body parts are preserved.

Syninclusion.

Crawler of Coccoidea (Fig. 1C, D).

Etymology.

The species is named in memory of our colleague coleopterist Prof. Aleksandr Vasilievich Puchkov.

Description.

Female. Habitus as in Figs 1B, 2C. Body not compact, slightly elongated and flattened. Body length 1.2 mm.

Coloration. Body, antenna, tegula, gaster dorsally and ventrally black; surface of frontovertex, mesoscutum, scutellum and axillae smooth, shiny, but without metallic shine, monotonously shallow reticulate with sparse punctures, evenly clothed in short setae (Figs 1B, 2A); coxae, and legs black, meso- and metatibia with lighter apices; protarsus dark; mesotibial spur, meso- and metatarsus light brown to yellowish.

Head. Hypognathous, slightly wider than thorax (8:7) in dorsal view, twice as broad as long, with rounded occipital margin; eyes bare, with inner orbits parallel; minimum distance between eyes almost 0.5 × head width; OOL about equal to posterior ocellar diameter (Fig. 2A); OCL about equal to half of posterior ocellar diameter; OOL:POL:LOL:OCL about 1:9:5.5:1.5; eye reaching occipital margin; facial cavity and interantennal prominence present but without distinct antennal scrobes (Fig. 2A); toruli located above of mouth margin, about at level of lower margin of eyes (Fig. 2B); malar sulcus complete; mandibles, probably two-toothed, with powerful teeth (Fig. 2B).

Antenna. Geniculate, 11-segmented (1:1:6:3); radicle short, about 2-2.5 × as long as broad; scape wide, flattened (Fig. 2A); pedicel conical, slightly shorter than F1-F4 combined, longer than any segment of funicle; funicle cylindrical, all segments broader than long, width of flagellomeres increases toward apex; F4-F6 and all segments of clava with mps (Fig. 2A, B); clava large, only slightly shorter than funicle, about 2.2 × as long as broad, with small oblique truncation on non-acute apical segment (Fig. 2B: ot), slightly wider than F6; flagellum and clava clothed in short setae.

Mesosoma. Pronotum short, almost vertical, in dorsal view without transverse dorsal surface (Fig. 2A); mesoscutum broader than long, flat; axillae transverse-triangular with anteromedial angles contiguous (Fig. 2A); prepectus large, bare, polygonally reticulate, posterior margin extended to base of tegula; acropleuron convex, bare, with shallow longitudinal reticulate sculpture, long, in lateral view twice as long as height (Fig. 2C: ac), with distinct acropleural sulcus; subtegular region (Fig. 2D: str) of acropleuron large, convex, delineated by distinct sulcus; metapleuron narrow, without visible setation (Fig. 2C).

Wings. Fully developed. Fore wing with basal cell uniformly setose; costal cell narrow; submarginal vein without distinct extension (Fig. 3A), although, when lighting from above, parastigma appears to be somewhat swollen (Fig. 1B: pst); hyaline break (unpigmented area) present; marginal vein long, but slightly shorter than stigmal and postmarginal veins (Fig. 1B, left wing); proportions of forewing venation as in Figs 1B, 2C, 3A; stigmal vein with long narrow uncus, consisting row of uncal sensillae (Fig. 3A: unc); enlarged seta marking apex of postmarginal vein of fore wing rather absent (Fig. 3A); setae of marginal fringe short.

Legs. Normal in size, alike polygonally reticulate; protibia with long, curved calcar; basitarsal comb poorly developed; tarsi 5-segmented; mesotibial spur slightly shorter than mesobasitarsus; metatibia with two spurs.

Gaster. As long as mesosoma, polygonal reticulate equal dorsally and ventrally, apical margins of metasomal terga straight, parallel; paratergites and cercal setae not visible; ovipositor stylet combined from 1st and 2nd valvulae (stylet suture presumably visible in Fig. 3C); hypopygium in Fig. 3B-D: hyp.

Male. Unknown.