Psechrus luangprabang Jäger, 2007

Figs 43a–d, 44a–e, 45a–e, 81d, 83b, 85l, 88m, 91m, 93a

Psechrus luangprabang Jäger 2007: 45, figs 48–53, 68–72 (Description of ♂ and ♀, illustration of ♂ and ♀). [Holotype ♂ (SB 1030) from LAOS: Luangprabang Province: Xieng Ngeun District, Ban Keng Koung [L15], N 19°40'58'', E 102°18'27'', 372 m, disturbed forest at riverside of Nam Khan, between rocks; P. Jäger & J. Altmann leg. 07.III.2006; SMF 56389; Paratypes: 1 ♂ (SB 1031), 2 ♀♀ (SB 1032, 1034) with same data as for holotype; SMF 56390(SB 1031–1032), SMF 56392(SB 1034); 1 ♂ (SB 1033) with same data as for holotype, but: leg. 08.III.2006; SMF 56391, all type material examined]. Bayer and Jäger 2010: 72.

Additional material examined (13 ♂♂, 24 ♀♀, 6 s.a. ♂♂, 10 s.a. ♀♀, 5 juvs). LAOS: Luang Nam Tha Province: Nam Ha Protected Area |2|, N 21°06'43'', E 101°20'36.1'', 693 m, under bridge; P. Jäger leg. 06.III.2008; 2 ♂♂ (SB 21–22), SMF (SB 21), MHNG (SB 22). Nam Ha Protected Area |3|, N 21°08'17.6'', E 101°24'07.3'', 746 m, under bridge; P. Jäger leg. 06.III .2008; 1 ♀ (SB 6), SMF . Nam Ha Protected Area |6|, N 21°03'32.1'', E 101°24'03'', 590 m, under bridges; P. Jäger leg. 07. III .2008; 3 ♂♂ (48, 53–54), 3 ♀♀ (SB 49, 51–52), 4 s.a. ♀♀ (SB 50, 56–58), SMF . Same data as above, but: P. Jäger & S. Bayer leg. 19.XI.2009; 1 ♂ (SB 459), 2 s.a. ♂♂ (SB 352, 359), SMF . Ban Tavan |3| (near Luang Nam Tha), N 20°58'57.6'', E 101°29'38'', 586 m, small stream with steep slopes at both sides, between roots and stones; P. Jäger & S. Bayer leg. 18.XI.2009; 1 ♂ (SB 402), 1 ♀ (SB508), 1 juv. (SB 356), SMF . Oudomxai Province: Lipi Provincial Protected Area, ca. 3.7 km S of Oudomxai, N 20°39'39.8'', E 101°59'47.3'', 741 m, under rock; L. Nophaseud leg. 18. IV.2011; 2 ♀♀ (SB 1143–1144), SMF . Luang Prabang Province: Nong Khiao, near Tham Pathok Cave, N 20°33'05'', E 102°37'55'', ca. 350 m, between rocks and stones; P. Jäger & S. Bayer leg. 16.XI.2009; 1 ♂ (SB 353), 1 s.a. ♀ (SB 346), 1 s.a. ♂ (SB 358), SMF . Luang Prabang, Phou Si, N 19°53'23'', E 102°08'04'', ca. 300 m, small, dry secondary forest, between stones; P. Jäger & S. Bayer leg. 14.XI.2009; 1 juv. (SB 360), SMF . Xieng Ngeun District, Ban Keng Koung [L15], N 19°40'58'', E 102°18'27'', 372 m, disturbed forest at riverside of Nam Khan, between rocks; P. Jäger & J. Altmann leg. 07. III.2006; 1 s.a. ♂ (SB 1035), SMF 56391 . Same data as above, but: P. Jäger leg. by night 23. II.2008; 2 ♀♀ (SB 41–42), SMF . Ban Ean, That Se [L 13], N 19°50'34'', E 102°13'07'', 304 m; P. Jäger & J. Altmann leg. 05. III.2006; 1 ♀ (SB 1036), SMF . Ban Pak Bak (at Nam Khan river), Houay Kho, N 19°44'09.1'', E 102°16'37.5'', 328 m, forest along stream; P. Jäger leg. by night 23. III.2007; 1 ♀ (SB 61), SMF . Ban Nong Di (at Nam Khan river), N 19°41'03.5'', E 102°21'31.2'', 280 m, forest along stream, between stones; P. Jäger leg. by night 21. III.2007; 1 ♂ (SB 67), 2 ♀♀ (SB 68–69), SMF . Vientiane Province: Ban Phoxay (near Vang Vieng), N 19°00'44'', E 102°26'46'', 260 m, in tunnel underneath the road; P. Jäger, M. Sandner & F. Steinmetz leg. 14. III.2007; 3 ♀♀ (SB 76, 91–92), SMF , except SB92 (MHNG). Same data as above, but: P. Jäger leg. 14.III.2008; 1 ♀ (SB 20), SMF . Same data as above, but: P. Jäger & S. Bayer leg. 12.XI.2009; 1 ♀ (SB 424), 1 s.a. ♀ (SB 362), 3 juvs (SB 332, 361, 363), SMF . Ban Don Makhay (near Vientiane), N 18°05'04'', E 102°40'15'', ca. 180 m, secondary forest; P. Jäger leg. by night 01. IV.2011; 2 ♂♂ (SB 997, 1001), 2 ♀♀ (SB 998, 1002), 1 s.a. ♀ (SB 1003), SMF , 1 ♂ (SB 999), 1 ♀ (SB 1000), CJW . Bolikhamsay Province: Nam Kading National Protected Area, Tad Vang Fong training centre, N 18°20'28.8'', E 104°08'37.5'', 150 m, disturbed secondary forest, in hole of a tree trunk; P. Jäger & L. Nophasead leg. 23. III.2011; 1 ♀ (SB 986), SMF . Lak Sao, N 18°13'38'', E 104°44'47'', ca. 530 m, between rocks, near cave entrance; P. Jäger & S. Bayer leg. 09.XI.2009; 1 s.a. ♀ (SB 375), SMF . Lak Sao, N 18°13'09.4'', E 104°56'36.9'', 556 m, rock wall; P. Jäger & S. Bayer leg. by night 08.XI.2009; 1 s.a. ♀ (SB 371), 1 s.a. ♂ (SB 342), SMF . Houay Khiao (W of Lak Sao), N 18°12'47.8'', E 104°49'51.5'', 500 m, paddy fields, between stones; P. Jäger & J. Martens leg. 04. III.2010; 2 ♀♀ (SB 516–517), SMF .

Revised diagnosis (see also diagnosis for himalayanus -group above). Males: Embolus base (EB) with particular flat, elongated, distally curved extension (distally means here the section directed to the proximal margin of T [tegulum]). T clearly broader than palpal tibia (Fig. 43b). Females similar to P. demiror sp. nov. in the shape of anterior part of epigyne, actually the anterior (distal) margins of lateral lobes (LL). These strongly sclerotised, semicircular and extending anterio-laterally (Fig. 44a, 45a, 47a, 88l–m). Distinguished by the distal endings of margins of LL pointing posteriorly and by the two distinctly developed longitudinal edges below copulatory openings (CO) (Figs 44a, 45a, 88m). Vulva similar to P.demiror sp. nov. Distinguished by the larger (in relation to copulatory duct) and differently shaped spermatheca (Fig. 44b, 45b).

Description. Male (measurements of holotype first, those of other specimens given as ranges in parentheses): Body and eye measurements. Carapace length 6.2 (5.7–8.4), carapace width 4.7 (4.1–6.1), anterior width of carapace 2.6 (2.2–3.2), opisthosoma length 8.8 (7.5–11.6), opisthosoma width 3.8 (2.8–4.0). Eyes: AME 0.32 (0.27–0.39), ALE 0.40 (0.35–0.44), PME 0.43 (0.37–0.47), PLE 0.42 (0.36–0.45), AME–AME 0.19 (0.17–0.20), AME–ALE 0.09 (0.04–0.09), PME–PME 0.22 (0.22–0.29), PME–PLE 0.30 (0.30–0.35), AME–PME 0.57 (0.49–0.58), ALE–PLE 0.43 (0.34–0.48), clypeus height at AME 0.77 (0.63–0.94), clypeus height at ALE 0.72 (0.58–0.92).

Cheliceral furrow with three promarginal and four retromarginal teeth.

Measurements of palp and legs. Leg formula: 1423. Palp 8.3 (7.2–10.0) [3.0 (2.8–3.6), 1.5 (1.2–1.7), 1.3 (1.2–1.7), 2.5 (2.0–3.0)], I 51.7 (46.0–66.8) [13.4 (11.5–17.2), 3.1 (2.6–4.1), 14.5 (13.0–19.0), 14.4 (12.8–18.9), 6.3 (5.8–7.6)], II 39.6 (35.5–51.6) [10.9 (9.7–14.0), 2.8 (2.2–3.5), 10.5 (9.4–14.0), 10.5 (9.4–14.4), 4.9 (4.7–5.7)], III 26.9 (24.0–35.5) [7.6 (7.0–10.4), 2.0 (1.8–2.6), 6.7 (5.7–8.8), 7.0 (6.2–9.4), 3.6 (3.3–4.3)], IV 42.1 (36.6–53.6) [11.2 (9.8–14.6), 2.4 (2.1–3.1), 10.5 (9.0–14.0), 12.2 (10.7–15.3), 5.8 (5.2–6.6)].

Spination. Palp: 131 (131), 110 (110), 1101 (1101); legs: femur I 566 (566), II 556 (566,567,667) III 555 (545), IV 554 (654); patella I–IV 000; tibia I–II 3038 (3038), III 3135{3034} (3136,3035), IV 3036 (3036,3037); metatarsus I–III 3035 (3035), IV 3036{3035} (3036,3035).

Palpal femur without modification, broadest distally (Fig. 43d).

Copulatory organ (see also diagnosis and general description for himalayanus -group). Conductor (C) and embolus (E) long, slightly S-shaped. T appears partly divided subdistally due to a curved ridge (Fig. 43b) and protruding quite distinctly out of cymbium alveolus (Figs 43a,c). Palpal tibia in lateral view medium sized (Fig. 43c). Cymbium dorsally with moderate dense scopula, covering 1/3 of cymbium (Fig. 83b).

Female:

Body and eye measurements. Carapace length 6.6–9.8, carapace width 4.4–7.3, anterior width of carapace 2.7–4.1, opisthosoma length 10.9–14.4, opisthosoma width 4.6–6.9. Eyes: AME 0.34–0.39, ALE 0.42–0.47, PME 0.42–0.48, PLE 0.42–0.47, AME–AME 0.21–0.32, AME–ALE 0.03–0.10, PME–PME 0.27–0.42, PME–PLE 0.37–0.53, AME–PME 0.58–0.83, ALE–PLE 0.42–0.67, clypeus height at AME 1.03–1.53, clypeus height at ALE 1.00–1.32.

Cheliceral furrow with three promarginal and four retromarginal teeth.

Measurements of palp and legs. Leg formula: 1423. Palp: 7.2–11.4 [2.5–3.9, 1.0–1.7, 1.3–2.1, 2.4–3.7]; Legs: I 33.8–54.4 [9.0–14.9, 2.6–4.1, 9.3–15.2, 8.6–14.0, 4.3–6.2], II 29.9–44.1 [8.3–12.5, 2.4–3.8, 8.1–11.8, 7.4–11.0, 3.7–5.0], III 21.2–32.2 [6.2–9.8, 1.9–3.0, 5.2–7.7, 5.2–7.9, 2.7–3.8], IV 29.9–46.3 [8.4–13.3, 2.1–3.3, 7.8–12.1, 7.8–11.9, 3.8–5.7]. Thus, legs in females, in relation to other Psechrus species, short: FEM-I+MTT-I/CL: 2.6–2.9.

Palpal claw with 13–16 teeth.

Spination. Palp: 131, 110, 1101, 1014; legs: femur I 566, II 566 (556,555), III–IV 555; patella I–IV 000; tibia I–II 3038, III 3036 (3035,3034), IV 3036; metatarsus I–III 3035, IV 3035 (3036).

Copulatory organ (see also diagnosis and general description of himalayanus -group). Median septum almost as long as broad (Fig. 44a). Slit sense organs and epigynal muscle sigilla outside epigynal field.

Primordial copulatory organ. Pre-epigyne: Pre-septum broader than long, with distal margins of pre-LL pointing posteriorly (Fig. 44e). Fine, transversal edges below pre-CO recognisable (Fig. 44e).

Pre-vulva: Pre-vulva similar to P. marsyandi . Distinguished by the longer pre-CD and the smaller pre-SH (in relation to pre-receptaculum) (Fig. 44d).

Colouration of male and female (see also description for himalayanus -group and Psechrus). Median bands on carapace serrated. Rarely with narrow, dark extensions connecting them with the lateral bands. Lateral bands narrow to medium sized (ca. 1–1.2 diameter of PME) and serrated. Light longitudinal line ventrally on opisthosoma may be continuous, broken subdistally or constricted subdistally and medium- sized. If measured centrally on opisthosoma, its width is less than the width of one half of the cribellum (ca. 0.5–0.7 of one half of the cribellum).

Variation of copulatory organs. Males: Without significant variation. Females: Anterio-lateral curves of distal margins of LL may be larger and extending further laterally (Fig. 45a, 88m). Main sections of margins of LL may be approximately straight (not illustrated). Vulva with less variation: Position and length of SH may differ slightly (Figs 44b, 45b, 91m) and margins of initial parts of CD may be less clearly visible (Fig. 45b). Transition zone between CD and spermatheca may differ in shape (Figs 44b, 45b, 91m).

Distribution. Laos (Fig. 98).