Psechrus pakawini sp. nov.

Figs 41a–h, 42a–e, 85k, 88j, 91j

Type material: Holotype ♂ (SB 140), THAILAND: Chiang Mai Province: Chiang Mai District, Doi Suthep, N 18°48'59'', E 98°53'34'', ca. 1600 m, mountainous rainforest; P. Schwendinger leg. 05.X.1993; MHNG ; Paratypes (4 ♂♂, 9 ♀♀, 1 s.a. ♀): 4 ♀♀ (SB 197, 199: SMF; SB 198, 200: MHNG), THAILAND: Chiang Mai Province: Chiang Mai District, Doi Suthep-Pui National Park, Doi Pui, San Koo, 1450 m, evergreen rainforest, along a small trail to the summit; P. Dankittipakul leg. 27.I.2001; SMF & MHNG (see above) ; 1 ♂ (SB 139), THAILAND: Chiang Mai Province: Fang District, Doi Angkhang, Ban Luang, ca. 1600 m; P. Schwendinger leg. 02.XI.1990; MHNG ; 1 ♂ (SB 142), THAILAND: Chiang Mai Province: Fang District, Doi Angkhang, ca. 1500 m; P. Schwendinger leg. 30.X.1987; MHNG ; 4 ♀♀ (SB 209–210: MHNG, SB 213–214: SMF), THAILAND: Chiang Mai Province: Fang District, Doi Angkhang, Royal Agricultural Research Station, ca. 1500 m, at a stream; P. Dankittipakul leg. 02.V.2002; MHNG & SMF (see above) ; 2 ♂ (SB 146–147), 1 ♀ (SB 148, actually subadult, but close to adult moult; adult epigyne already fully developed; could be dissected and examined), THAILAND: Chiang Mai Province: Chomthong District, Doi Inthanon, ca. 1700 m, rain forest; P. Schwendinger leg. 08.X.1987; MHNG .

Additional doubtful material examined. MYANMAR: Kachin Province: Namti, Loglai; 1945; Coll. J.E.S.; 1 ♀ (SB 974), CAS 9032227 .

Etymology. The specific name is a patronym in honour of the collector of part of the type material, Pakawin Dankittipakul; noun (first name) in genitive case.

Diagnosis (see also diagnosis for himalayanus -group above). Males similar to P. ghecuanus in having a quite strongly sclerotised, semicircular extension apically on tegulum (T) (see asterisk in Fig. 39g). Distinguished by the longer embolus (E) and conductor (C) (the former with proximal section depress, in ventral view) (Figs 41a–c). Moreover, E arising proximal to the arising point of C (Fig. 41b) or at the same level on T. Females similar to P. ghecuanus in shape of epigyne and vulva. Distinguished by the margins of lateral lobes (LL) almost 180° curved anteriorly and pointing posteriorly (Figs 41e, 42c,e) and by the longer copulatory ducts (should be viewed from frontal, Fig. 41h) extending further medially.

Description. Male (measurements of holotype —largest male— first, those of paratype SB 139 —smallest— in parentheses):

Body and eye measurements. Carapace length 6.8 (3.9), carapace width 4.9 (2.8), anterior width of carapace 2.6 (1.6), opisthosoma length 7.8 (4.6), opisthosoma width 3.0 (1.7). Eyes: AME 0.35 (0.24), ALE 0.43 (0.30), PME 0.46 (0.32), PLE 0.44 (0.28), AME–AME 0.14 (0.12), AME–ALE 0.03 (0.04), PME–PME 0.16 (0.14), PME–PLE 0.28 (0.22), AME–PME 0.47 (0.38), ALE–PLE 0.34 (0.23), clypeus height at AME 0.68 (0.47), clypeus height at ALE 0.63 (0.46).

Cheliceral furrow with three promarginal and four retromarginal teeth.

Measurements of palp and legs. Leg formula: 1423. Palp 8.2 (5.2) [3.0 (1.9), 1.3 (0.8), 1.2 (0.8), 2.7 (1.7)], I 51.9 (38.1) [13.5 (10.5), 3.2 (1.9), 14.4 (10.6), 14.4 (10.2), 6.4 (4.9)], II 42.0 (28.0) [11.6 (9.0), 2.7 (1.4), 11.2 (7.7), 11.5 (7.0), 5.0 (3.9)], III 29.0 (19.5) [8.5 (5.7), 2.2 (1.3), 7.1 (4.9), 7.6 (4.9), 3.6 (2.7)], IV 42.2 (30.0) [12.1 (8.3), 2.5 (1.4), 10.4 (7.6), 11.7 (8.3), 5.5 (4.4)].

Spination. Palp: 131 (131), 110 (110, in both specimens prolateral spine very small), 1101 (1101, in both specimens all spines very small); legs: femur I 556 (556), II 556 (546) III 555 (545), IV 544 (545); patella I–IV 0000; tibia I 3038 (3038), II 3038 (4048), III–IV 3036 (3134); metatarsus I–III 3035 (3035), IV 3036 (3035).

Palpal femur without modification, broadest distally (Fig. 41d).

Copulatory organ (see also diagnosis and general description for himalayanus -group). Conductor fleshy, straight and distally in retrolateral section with longitudinal bending (Figs 41b–c). Sperm duct extending quite far proximally at T. Palpal tibia in lateral view short (Fig. 41c). Cymbium dorsally without scopula.

Female: Body and eye measurements. Carapace length 6.0–8.4, carapace width 4.3–6.1, anterior width of carapace 2.7–3.7, opisthosoma length 7.9–11.8, opisthosoma width 4.3–6.1. Eyes: AME 0.33–0.41, ALE 0.42–0.45, PME 0.46–0.48, PLE 0.41–0.48, AME–AME 0.18–0.26, AME–ALE 0.04–0.06, PME–PME 0.25–0.30, PME–PLE 0.36–0.39, AME–PME 0.48–0.63, ALE–PLE 0.41–0.51, clypeus height at AME 0.76–0.99, clypeus height at ALE 0.71–0.98.

Cheliceral furrow with three promarginal and four retromarginal teeth.

Measurements of palp and legs. Leg formula: 1423. Palp: 7.2–9.5 [2.3–3.4, 1.2–1.4, 1.3–1.6, 2.4–3.1]; Legs: I 33.7–45.7 [9.2–12.5, 2.6–3.5, 9.4–12.7, 8.2–11.7, 4.3–5.3], II 27.7–36.8 [7.9–10.2, 2.3–3.2, 7.3–9.7, 6.6–9.4, 3.6–4.3], III 19.4–26.1 [5.9–7.9, 1.7–2.4, 4.4–6.2, 4.8–6.4, 2.6–3.2], IV 27.7–37.2 [7.9–10.5, 2.0–3.0, 7.0–9.5, 6.9–9.6, 3.9–4.6]. Thus, legs in females, in relation to other Psechrus species, short: FEM-I+MTT-I/CL: 2.85–2.9.

Palpal claw with 12–14 teeth.

Spination. Palp: 131, 110, 1101, 1014; legs: femur I 566 (546), II 546 (545), III 545 (535), IV 544; patella I–IV 000; tibia I–II 3038, III 3034 (2024), IV 3036; metatarsus I–III 3035, IV 3035 (3033).

Copulatory organ (see also diagnosis and general description of himalayanus -group). Slit sense organs (SO) and epigynal muscle sigilla (EM) outside epigynal field (EF). Anterior to epigyne many slightly curved wrinkles (Fig. 41e).

Primordial copulatory organ. Pre-epigyne: Similar to P. ghecuanus . Distinguished by the absence of a pre-EF and by two fine ridges medially (Fig. 42a).

Pre-vulva: Similar to P. ghecuanus . Distinguished by the less developed pre-SH (Fig. 42b). It is difficult to discriminate pre-SH and pre-receptaculum in the s.a. ♀ of this species.

Colouration of male and female (see also description for himalayanus -group and Psechrus). Median bands on carapace serrated. Lateral bands narrow (at most 2/3 diameter of PME) and serrated. Light longitudinal line ventrally on opisthosoma may be continuous, broken subdistally or constricted subdistally and medium-sized. If measured centrally on opisthosoma, its width is less than the width of one half of the cribellum (ca. 0.5–0.8 of one half of the cribellum).

Variation of copulatory organs. In males E and C may be a bit longer (Fig. 85k) than in holotype (Fig. 41b). Females: Distal parts of margins of LL extending differently far posteriorly (Figs 41e, 42c,e, 88j). MS in a few specimens slightly shorter (Fig. 42c). In vulva SH may be longer (Fig. 41f, 91j) or shorter (Fig. 42d).

Remark: The female specimen SB 974 from Kachin Prov., Myanmar slightly differs from the type specimens. Its spermathecae have a rather inhomogeneous shape (not illustrated). With further material from this region, especially males, it may be possible to provide a definite identification.

Distribution. Thailand (Fig. 98).