Barinas guanenta sp. nov.

Figs 1, 2, 4

Type material.

Colombia• ♂ holotype: Santander, Cabrera, Cueva del Indio de Doña Joaquina; [6.562261; - 73.237897]; [1100 m]; 14 July 2007; Ca-vita grupo de bioespeleologia leg.; ICN-Ao-742.1 • 1 ♂ 1 ♀ paratypes; same data as the holotype; ICN-Ao-742.

Etymology.

Guanentá was the leader of the Guanes, an indigenous group that fought bravely against the Spanish conquerors in the region where the species was collected. Noun in apposition.

Diagnosis.

Can be distinguished by having pedipalps entirely pale yellow without variegated pattern (in B. piragua distal portion of Tr and Fe black, and tarsi entirely black; in B. virginis comb. nov. with a variegated pattern on Fe-Ti); variegated coloration on areas I-IV ( B. flava and B. piragua without variegated coloration); stylus of penis straight (in all other Barinas species the stylus is curved at apical portion) (Figs 1, 2).

Description.

(Figs 1, 2). Male holotype (ICN-Ao-742.1) measurements (mm): CL (0.57), AL (1.22), CW (1.20), AW (1.44), DSL (1.79), IOD (0.27), ChL (0.86). Pedipalp: Tr (0.35), Fe (0.48), Pa (0.34), Ti (0.57), Ta (0.44), Claw (0.42). Legs I-IV (Tr/Fe/Pa/Ti/Mt/Ta/TL): I (0.21/2.45/0.43/1.93/3.69/1.24/9.95), II (0.27/6.19/0.65/5.38/7.30/3.57/23.36), III (0.34/4.02/0.75/2.47/4.85/1.29/13.72), IV (0.37/6.11/0.71/3.57/6.80/1.79/19.35). Dorsum: DS outline Epsilon type 2 (Figs 1A, B, 2A). Cheliceral sockets narrow, separated by medial projection (Fig. 2A). Anterior margin of DS smooth. Ocularium high (Figs 1D, E, 2B). Carapace smooth. Mesotergum delimited, divided into four areas (area I divided into two halves), with a transverse row of granules on each (Figs 1B, D, 2A, B). Areas I-IV with two paramedian spines, AHF: I = IV <II = III (Figs 1D, E, 2B); paramedian spines of areas III and IV projected backward (Figs 1D, E, 2B). Posterior margin of DS substraight, with a row of tubercles. Free tergites I-III with a row of tubercles and anal operculum with some tubercles (Figs 1B, 2A, B). Venter: Tegument granular. Coxa I with one trifid tubercle on the anterior margin and a transverse row of tubercles on the medial region (Fig. 2C). Coxa III-IV with some tubercles. Sternites with rows of spaced granules (Fig. 1C, D). Stigmatic area slightly granular. Stigmata large and oval (Fig. 1C). Genital operculum slightly granular. Chelicerae: Chelicera swollen. Segment I rectangular in dorsal view, with three-four tubercles on the proximal region and one-two tubercles on the ectal face (Figs 1B, 2A). Basal region of the movable finger with abundant setiferous tubercles of different sizes (some reaching the medial region of hand) (Fig. 2D). Fixed finger with smooth inner surface (Fig. 2D). Movable finger with one trapezoid sub-basal tooth and distal inner surface dentate (Fig. 2D). Pedipalps: Trochanter with five dorso-basal tubercles and two ventroectal tubercles. Fe with a ventroectal row of four setiferous tubercles (the two basalmost largest and the two distalmost small-sized), and one large ventromesal setiferous tubercle in the distal portion. Pa armed with one large ventromesal setiferous tubercle (Fig. 2E). Tibial setation: ectal iiIi, mesal IIi. Ta setation: ectal I?i, mesal IIi. Legs: Leg I-IV granular. Leg I filiform, the rest getting steadily thicker from leg II-IV (Fig. 1A). Tr I-III slightly granular, Tr IV with two dorsodistal tubercles (Figs 1B, 2A). Tarsal counts: 7(3)-7(3)/17(3)-16(3)/7-7/8-7. Penis: LP small and depressed, crescent-shaped, with anterolateral short sharp corners, apically pointed (Fig. 2F). Malleus with two pairs of trifid MS-A, and one pair of trifid MS-B (Fig. 2F); MS-C absent. Two pairs of MS-D located in a keel between the ventral part of LP and the base of the stylus; MS-E2 large and trifid, MS-E1 medium-sized (half of the size of MS-E2) (Fig. 2F). Stylus straight and elongated, surpassing the LP, with a short DP at the proximal region, and a DH just after the union of the DP with the stylus (Fig. 2G, F). Color (in ethanol) (Fig. 1A-E): Carapace and mesotergum reticulated Dark Grayish Olive (111) on Light Grayish Olive (109). Pedipalps and chelicerae Yellowish White (92). Free tergites Dark Olive (108). Coxae I-IV Light Grayish Olive (109) with lateral margins Dark Olive (108), trochanters I-IV Light Grayish Olive (109). Fe, Pa, and Ti I Dark Olive (108). Femora and tibiae II-IV Light Grayish Olive (109), patellae II-IV Dark Olive (109).

Female paratype (ICN-Ao-742). Similar to male, except for the non-hipertelic chelicera and shorter Fe IV. Measurements (mm): CL (0.57), AL (0.93), CW (1.06), AW (1.24), DSL (1.50), IOD (0.23), ChL (0.61). Pedipalp: Tr (0.22), Fe (0.49), Pa (0.32), Ti (0.46), Ta (0.43), Claw (0.33). Legs (Tr/Fe/Pa/Ti/Mt/Ta/TL): I (0.26/2.00/0.42/1.52/2.96/1.05/8.21), II (0.34/4.87/0.50/4.54/6.08/3.69/20.02), III (0.31/3.70/0.64/2.21/4.02/1.16/12.04), IV (0.34/5.52/0.65/?/?/?/?). Tarsal counts: 7-7/17-?/7-?/?-?.

Distribution.

Known only from the type locality, a cave in the Magdalena Valley montane forest ecoregion (NT0136) (Fig. 4).