Neosisyphus quadricollis (Gory, 1833) (Figs. 26, 27A)

Montreuil 2015b: 20.

Sisyphus quadricollis Gory, 1833

Gory 1833: 9; Harold 1869: 985; Péringuey 1901: 87; Arrow 1927: 459; Haaf 1955: 345; Ferreira 1972: 829.

Type locality: Cape of Good Hope.

Size: Male: length: 13.5–11; width: 6.5–5.4; female: length: 13.8–11; width: 6.5–5.5.

Diagnosis: N. quadricollis is close in appearance to N. kuehni . However, N. quadricollis may be separated by a combination of the following characters: elytra emarginated laterally with the concavity of the margin distinctly deep in the middle; the projecting metatrochanter in males is one-third of the total length of the metafemur; the metafemur lacks spines.

Examined non-type material: See Supplementary information.

Distribution: N. quadricollis has been recorded from deep sands and natural shrubland under drier, warmer winter rainfall climate on the southwest coast of South Africa (Western Cape) (Davis 1987, 1997; Davis et. al. 2008). In this region, it is active for a short period between late October to late December. It also has been collected between November and March at Willowmore and Lemoenkloof (Eastern Cape) in karoo vegetation of the late summer rainfall region (Fig. 28).