Sisyphus costatus (Thunberg, 1818) (Figs. 11, 12B).

Peringuéy 1901:100; Gillet 1912:22; Arrow 1927:458; Haaf 1955:355–6; Ferreira 1972: 794–5; Harold 1869:984

Ateuchus costatus Thunberg, 1818

Thunberg 1818:412

Size: Male: length: 5.3– 4.8 mm, width: 3.0–2.0 mm. Female: length: 5.3– 4.2 mm, width: 2.6– 2 mm.

Type locality: (Cape of Good Hope) South Africa

Diagnosis: S. costatus is close in appearance to S. australis sp. n. In both species, the elytral interstriae bear alternating rows of well-developed or less-developed setae (Fig. 11F). However, S. costatus is distinguished by an upcurved anterior clypeal edge and a concavity between the medial clypeal teeth. In addition, it lacks a carina between the clypeus and the frons (Fig. 11G).

Examined type material

(1♀ SMNH) Lectotype (designated here): Red label / Uppsala University Zoology Museum; Thunberg, saml.nr. 3206., Ateuchus costatus, Cap. minutus. F. TYP/ / Lectotype, det. GM Daniel, 2016 /.

Examined non-type material: See Supplementary information.

Distribution: S. costatus has been recorded in moist upland savanna and highland grassland (Davis et al. 1999, Davis et al. 2005) from the Eastern Cape to the north of South Africa (Fig. 13).

Remarks: The species was described by Thunberg (1818) as Ateuchus costatus . Gillet (1912) transferred Thunberg’s species to the genus Sisyphus .