Empria tridentis Lee & Ryu, 1996

Figs 1C, D, F, H, K, L, 6B, 7B

Empria tridentis Lee & Ryu, 1996: 23.

Diagnosis.

Female (Figs 1C, F, K, 6B). Body length 5.8-6.1 mm. Head and thorax black (clypeus pale in lower margin), except labrum, posterodorsal margin of pronotum, tegula, apex of all femora, basal 1/3 of fore and middle tibiae, basal 2/3 (mostly) of hind tibia, basal half of middle and hind tarsomere 1 white (pale); cenchrus, all trochanters and trochantelli yellowish brown; apical 2/3 of fore tibia, fore tarsus and middle tibia anteriorly brown; mandible at apex reddish dark brown; labial and maxillary palps dark brown; abdomen dark brown, abdominal terga with 3-4 pairs of whitish (pale) patches (the tergum 5 with small-sized or indistinct patch) (Fig. 1C, F, K). Malar space 1.7-1.9 times as long as diameter of median ocellus (Fig. 1K). Flagellum 2.3-2.5 times as long as head width. Ovipositor sheath only reaching beyond apex of abdomen; dorsal and ventral margins slightly convergent and apex obliquely truncated (Fig. 1F). Lancet with about 17 serrulae, membrane between serrulae weakly convex; each serrula subtriangularly protruding basally (Fig. 6B), with a basal denticle and 2-7 distal denticles.

Male (Figs 1D, H, L, 7B). Body length 4.7-5.1 mm. Similar to female, but fore leg much paler than female; middle tibia anteriorly and hind tibia at base white (pale); middle and hind tarsomere 1 entirely dark brown; sometimes abdominal segments with narrow posterior whitish margins (Fig. 1D, H, L). Malar space 1.4-1.6 times as long as diameter of median ocellus (Fig. 1L). Flagellum 3.6-3.7 times as long as head width. Posterior margin of sternum 9 rounded. Penis valve as in Fig. 7B, valviceps almost equal in length to valvura.

Genetic data.

Based on the barcode region of two available COI sequences (Japan), the distance between them is 0.2%. The nearest neighbours are species of the E. longicornis group, diverging by a minimum of 4.9%. Based on the nuclear data of one specimen (Japan), the nearest neighbours are species of the E. longicornis group, diverging by a minimum of 2.4%.

Type specimens examined.

South Korea: ♀, holotype of Empria tridentis (YNU-Sym-0011), " Gangwon-do, Goseong-gun, Hyangnobong, 38°19'N, 128°18'E, 13.V.1992, S.M. Ryu " (YNU) ; 8♀♀, paratypes of Empria tridentis (YNU-Sym-1115~1121, 1123), "same data as holotype" (1♀ in CSCS, 2♀♀ in NIBR, 5♀♀ in YNU) .

Additional specimens examined.

South Korea: 1♂, Gangwon-do, Pyeongchang-gun, Mt. Balwangsan, 11.VI.2000, J.W. Lee (YNU) ; 1♀ 4♂♂, Gangwon-do, Wonju-si, Mt. Baegunsan, 37°15'30.5"N, 127°58'55.11"E, 19.V-6.VI.2011, H.Y. Han (YNU) ; 2♂♂, same data, J.W. Lee (YNU); 2♂♂, Gangwon-do, Wonju-si, Panbu-myeon, Mt. Baegunsan, 28.IV-24.V.2012, H.Y. Han (YNU) ; 1♀ 1♂, Jeollanam-do, Jangseong-gun, Bukha-myeon, Hyangnobong, 37°27'00"N, 126°51'00"E, 13.V.1992, J.W. Lee (YNU) ; 3♀♀ 1♂, same locality, 13.VI.1992, J.W. Lee (1♀ 1♂ in NSMT, 2♀♀ in YNU) .

Host plants.

Rosaceae: Filipendula camtschatica (Pall.) Maxim., Geum japonicum (Shinohara et al. 2015).

Distribution.

Japan, Russia (Prous et al. 2011a), South Korea (Lee and Ryu 1996).