Guapinannus clava, sp. n.

(Figs 2, 5–7, 9)

Holotype. Male. PANAMA: Bocas del Toro: Almirante, at dam on Nigua Creek, 9.25°N 82.65°W, 810m, 23 Mar 1959, H. S. Dybas, 1♂ (UCR _ ENT 00097823) (FMNH).

Paratypes: COSTA RICA: Puntarenas: 5 km SW Finca Las Cruces, La Fila, 8.78937°N 82.98133°W, 1433 m, 21 Mar 1973, J. A. Wagner, J. B. Kethley, 1♂ (UCR _ ENT 00090603) (FMNH) . PANAMA: Bocas del Toro: Almirante, at dam on Nigua Creek, 9.25°N 82.65°W, 1080m, 23 Mar 1959, H. S. Dybas, 2♂ (UCR _ ENT 00090784, UCR_ENT 00097822), 7♀ (UCR _ ENT 0009782, UCR_ENT 00097824-UCR_ENT 00097829) (FMNH) ; 25 Mar 1959, H. S. Dybas, 1♂ (UCR _ ENT 00090740) (FMNH) . Almirante, trail to dam on Nigua Creek, 9.3003°N 82.40214°W, 5 m, 23 Mar 1959, H. S. Dybas, 3♀ (UCR _ ENT 00090748,UCR_ENT 00090778, UCR_ENT 00090779), 4 ; u (UCR _ ENT 00090757, UCR_ENT 00090758, UCR_ENT 00090760, UCR_ENT 00090761), 2♂ (UCR _ ENT 00090756,

UCR_ENT 00090777) (FMNH). Bocas del Toro: Almirante, trail to dam on Nigua Creek, 9.29792°N 82.4211°W, 15m, 31 Mar 1959, H. S. Dybas, 2♂ (UCR _ ENT 00090715, UCR_ENT 00090717) (FMNH) .

Etymology: Named for the club-shaped vesica after “clava” Latin for club (noun in apposition).

Diagnosis: Recognized by the mid-sized body length (1.54 mm), oval forewing outline (Fig. 2), fairly broad C+Sc (black arrow), rc (black star) and rc1 relatively short and wide, large vein-tracing areoles associated with scc, rc1, and rc (white arrow) (Fig. 6A), vesica short and thick with thick, rounded, club-shaped tip, and long, thin, and curved anophoric process (Figs 6F, 7).

Description: Male: Total body length 1.54 mm; length from posterior margin of pronotum to posterior wing margin 1.16 mm; greatest width across pronotum 0.66 mm; greatest width across forewings 0.87 mm. General coloration: mix of pale and darker brown or reddish brown. Vestiture: Dorsum with sparse setae, setae short; setae on posterior margin of forewing long. Structure: Head: Eye almost half as high as head; muscle scars large.

Thorax: Anterior pronotal margin concave, posterior margin slightly concave; pits on pronotum medium-sized, evenly distributed; scutellum in dorsal view abruptly narrowed to tip, tip pointed. Forewing: With outline broad anteriorly, oval; membrane contributing about one third of forewing length, posterior distal margin squarish, veintracing areoles large, distributed along scc, rc1, and rc; C+Sc fairly broad, wider at widest part of wing; distal process of R pronounced; An1 wing organ on corium weakly inflated, rounded, with median notch, without notch articulating with claval process; rc1 about 3 times as long as wide, much wider anteriorly; rc about 4 times as long as wide; tc about 3 times as long as wide; part of An1 and Cu forming posterodistal margin of tc slightly wider than other veins distal margin of rc1, rc, and tc cells double s-shaped; rc2–3 trapezoidal; cub almost reaching wing margin; dc1 horse head-shaped. Legs: Caws long. Genitalia (Fig. 7): Pygophore without lobe on right side; vesica with about half loop, not reaching right pygophore margin, thick at midpoint, with blunt apex, close to apex rounded; anophoric ridge with process, process long, with single lobe curved, tip pointed; right paramere curved, tapering, basal process broad; left paramere straight, tapering, basal process narrow.

Female: As in male and generic description.

Collecting method and habitat: Collected using Berlese extraction, Winkler extraction in fibrous center of decaying palm log.