Oeonus Godman, 1900, new status, is a subgenus of Oxynthes Godman, 1900

Oeonus Godman, 1900 -XI (type species Oeonus pyste Godman, 1900) clusters closely with Oxynthes Godman, 1900 -X (type species Goniloba corusca Herrich-Schäffer, 1869) in the genomic tree (Fig. 4 blue). COI barcode difference between the type species of these two genera is 9.6% (63 bp). Genetic differentiation between Oeonus and Oxynthes is approximately the same as between Orthos orthos and Orthos minka that were formerly considered conspecific (Fig. 4 magenta and green). Both Oeonus and Oxynthes include a small number of species and are not separated from each other by a prominent break (i.e, longer internal branches) comparable to that separating them both from Testia Grishin, 2019 . Therefore, we propose to treat these species as congeneric and place Oeonus, stat. nov. as a subgenus of Oxynthes .