Ambanus Ovtchinnikov, 1999: 63

WANG, XIN-PING, 2002, A Generic-Level Revision Of The Spider Subfamily Coelotinae (Araneae, Amaurobiidae), Bulletin of the American Museum of Natural History 2002 (269), pp. 1-150 : 26-31

Ambanus Ovtchinnikov, 1999: 63 .

TYPE SPECIES: Ambanus mandzhuricus Ovtchinnikov, 1999 , by original designation.

DIAGNOSIS: Females resemble Robusticoelotes in having a large epigynal atrium, broad copulatory ducts, and in lacking epigynal teeth, but they have short, posteriorly situated spermathecae (figs. 57, 58). Males lack a patellar apophysis and have a modified lateral tibial apophysis (figs. 59–61). Both sexes have only three promarginal and two retromarginal cheliceral teeth (fig. 63), unlike those of Robusticoelotes , which have more than five promarginal and retromarginal cheliceral teeth.

DESCRIPTION: See description of type species (below).

DISTRIBUTION: Northeastern China, far eastern Russia, Japan, Korea (map 1).

COMPOSITION: Eighteen species are includ­ ed, including 6 new combinations.

1. Ambanus amurensis Ovtchinnikov, 1999 : female holotype from Khaborovsk, far eastern Russia, in CSO, not examined.

2. Ambanus bifidus ( Paik, 1976) : female holotype from Mt. Kaya, Korea, in Kyungpook National University, Korea, not examined. NEW COMBINATION (transferred here from Coelotes ).

3. Ambanus circinalis (Gao et al., 1993) : One male paratype from Liaoning, China, in IZB, examined. NEW COMBINATION (transferred here from Coelotes ).

4. Ambanus dimidiatus (Paik, 1974) : male holotype, male and female paratypes from Mt. Pargong, Dongwha­sa, Korea, in Kyungpook National University, Korea, not examined.

5. Ambanus euini ( Paik, 1976) : female holotype and paratypes from Mt. Ilwol, Korea, in Kyungpook National University, Korea, not examined.

6. Ambanus grandivulvus (Yaginuma, 1969) : female holotype from Mt. Mitake, Tsushima, Nagasaki, in NSMT, examined.

7. Ambanus kayasanensis (Paik, 1972) : male holotype from Korea, in Kyungpook National University, Korea, not examined.

8. Ambanus kimi (Paik, 1974) : male ho­ lotype and paratypes from Mt. Kaya and Mt. Pargong, Korea, in Kyungpook National University, Korea, not examined.

9. Ambanus lunatus ( Paik, 1976) : female holotype and paratypes from Mt. Sokkri and Kwangreung, Korea, in Kyungpook National University, Korea, not examined. One female specimen from Korea, in KAI, examined.

10. Ambanus mandzhuricus Ovtchinnikov, 1999 : male and female paratypes from Kedrovaya Pad Reservation, S­Primorie, far eastern Russia, in SZM, examined.

11. Ambanus meniscatus (Zhu and Wang, 1991) : female holotype and paratypes from Tonghua, Jilin, China, in NBUMS, not examined. NEW COMBINATION (transferred here from Coelotes ).

12. Ambanus napolovi Ovtchinnikov, 1999 : male and female paratypes from Jasnoje viu, Przhewalsky Mts. East end, S­Primorie, far eastern Russia, in AMNH, examined).

13. Ambanus ovatus ( Paik, 1976) : female holotype from Mt. So­Paik, Korea, in Kyungpook National University, Korea, not examined.

14. Ambanus paiki Ovtchinnikov, 1999 : male and female paratypes from far eastern Russia, in SZM, examined.

15. Ambanus paikwunensis (Kim and Jung, 1993) : male holotype and female paratype from Mt. Paikwunsan Valley, Ildongmyon, Pochon­gun, Korea, in KAI, not examined. NEW COMBINATION (transferred here from Coelotes ).

16. Ambanus quadrativulvus (Paik, 1974) : male holotype, male and female paratypes from Paikrok­dam, Mt. Hanra, Quelpart Isl., Korea, in Kyungpook National University, Korea, not examined. Male and female specimens from Cheju­do, Korea, in KAI, examined.

17. Ambanus rostratus ( Song et al., 1993) : male paratype from Huanren, Liaoning, China, in IZB, examined. NEW COMBINATION (transferred here from Coelotes ). One female specimen from Liaoning, China, in IZB, examined.

18. Ambanus terdecimus ( Paik, 1978) : female holotype from Waheyonri, Geoje­do, Kyungnam, Korea, in Kyungpook National University, Korea, not examined. NEW COM­ BINATION (transferred here from Coelotes ).

Ambanus mandzhuricus Ovtchinnikov Figures 57–63

Ambanus mandzhuricus Ovtchinnikov, 1999: 63 , figs. 1–5 (2 male and 2 female paratypes from Kedrovaya Pad Reservation, S­Primorie, far eastern Russia, in SZM, examined).

DIAGNOSIS: This species is similar to A. paiki Ovtchinnikov but can be distinguished by the shape of copulatory ducts and the dorsally situated spermathecal heads of females (fig. 58) and by the different shape of the embolus and the broad conductor of males (figs. 59–61).

DESCRIPTION: Total length 10.0 –12.0. From front, anterior eye row slightly procurved, posterior row procurved; eye sizes and arrangements: AME smallest, about half ALE size or slightly larger, ALE subequal to PLE or slightly larger, PME slightly smaller than PLE; AME­AME subequal to AME­ ALE, ALE­PLE close together, PLE­PME roughly PLE diameter, PME­PME roughly AME diameter (fig. 62). Clypeal height slightly more than twice AME diameter; chilum present, divided, elongated. Chelicerae with three promarginal and two retromarginal teeth (fig. 63). Labium slightly longer than wide. Length of female 1st leg patella + tibia shorter than carapace length.

Tarsal organ and trichobothria not examined. Tarsal organ of Ambanus napolovi (from far eastern Russia) situated close to distal end of tarsus, anterior of distalmost trichobothrium (figs. 67–69).

Spinnerets not examined. Female spinnerets of A. napolovi : apex of ALS with 2 major ampullate gland spigots (MAP), 20 piriform gland spigots; PMS with at least 57 aciniform gland spigots, 2 cylindrical gland spigots (CY), and 1 or 2 minor ampullate gland spigots (mAP); PLE with at least 78 aciniform gland spigots and 3 cylindrical gland spigots (figs. 64–66). Spinnerets of A. rostratus also showed 3 cylindrical gland spigots on female PLS.

Female epigynum without epigynal teeth; atrium large, deep; copulatory ducts broad; spermathecal heads small, situated laterad of spermathecae; spermathecae short, slightly convoluted, with indistinct stalks and bases (figs. 57, 58). Male palp with patellar apophysis absent; RTA moderately long, with dis­ conductor lamella small; embolus basal in origin, strong, long, with spiral apex; median apophysis reduced (figs. 59–61).

MATERIAL EXAMINED: RUSSIA: Far East : S­Primorie, ‘‘ Kedrovaya Pad’ ’ Reservation, August 31, 1978, 1 male and 1 female paratype (D. Sherbakov, SZM) ; S­Primorie, ‘‘ Kedrovaya Pad’ ’ Reservation, June 27, 1976, 1 male and 1 female paratype (B. P. Zakharov, SZM) .

tal end slightly extended beyond tibia; lateral tibial apophysis modified to small concavity on distal tibia; cymbial furrow short, shallow, with dorsal edge not apparent; tegular sclerite more or less unsclerotized; conductor broad; conductor dorsal apophysis small;

Figures

Figs. 57, 58. Ambanus mandzhuricus Ovtchinnikov, female. 57. Epigynum. 58. Vulva.

Figs. 59, 60. Ambanus mandzhuricus Ovtchinnikov, male palp. 59. Ventral view. 60. Retrolateral view.

Figs. 61–63. Ambanus mandzhuricus Ovtchinnikov. 61. Male palp, prolateral view. 62. Eyes, dorsal view. 63. Chelicera, ventral view.