Philonthus madianus Bernhauer, 1936

Hromádka, Lubomír, 2008, Revision of Afrotropical species of the Philonthus peripateticus species group (Coleoptera: Staphylinidae: Philonthina), Acta Entomologica Musei Nationalis Pragae 48 (1), pp. 51-65 : 59-61

Philonthus madianus Bernhauer, 1936

( Figs. 25-28 )

Philonthus (Philonthus) madianus Bernhauer, 1936: 263 .

Type locality. Angola, Ebanga.

Type material examined. HOLOTYPE: ♁, ‘ Angola, Miss. Sc. Suisse, 1932-1933 / Ebanga nov., / Chicago NHMus., M. Bernhauer Collection. // madianus Bernhauer , Typus. [ochre oblong label, handwritten]’ ( FMNH) . PARATYPE: ♀, ‘ UGANDA, Madi, v.1927, / G. D. H. Carpenter, / Brit. Mus. Don. Marshall, / Chicago NHMus. M. Bernhauer Collection, // madianus Bernh. Cotypus. [ochre oblong label, handwritten]’ ( FMNH).

Redescription. Body length 11.9-12.2 mm (with stretched abdomen), length of fore body (to the end of elytra) 5.2-5.5 mm.

Colouration. Black, only clypeus along its anterior margin and antennal sockets narrowly yellow-brown; base of antennomere 2 and femora brown.

Head distinctly transverse, wider than long (ratio 39: 36 in male, 39: 33 in female), with well marked posterior angles, subparallel-sided, only medially slightly concave. Eyes almost as long as temples (ratio 14: 15). Four coarse punctueres between eyes, distance between medial interocular punctures about four times as large as distance between medial and lateral interocular punctures. Temporal area with many punctures of different size and with one long black bristle. Surface without microsculpture.

Antennae long, slightly widened towards apex, reaching posterior fifth of pronotum when reclined. Antennomere 1 distinctly shorter than antennomeres 2-3 combined, antennomere 2 slightly shorter than antennomere 3, antennomeres 4-10 of equal length, antennomere 11 somewhat longer than antennomere 10.

Pronotum as long as wide, subparallel-sided, anterior angles almost rectangular, with very apex obtusely rounded, posterior angles markedly rounded. Each dorsal row with two puntures, far away from each other; sublateral rows missing. Surface without microsculpture.

Scutellum coarsely and densely punctate, punctures somewhat smaller than those of elytra. Setation black.

Elytra distinctly wider than long (ratio 61: 55), parallel-sided, vaguely widened posteriad, posterior margin relatively deeply cut out. Punctation coarse and dense, punctures somewhat larger than eye-facets, interspaces smaller than punctures. Surface without microsculpture, setation grey.

Legs. Metatibia as long as metatarsus. Metatarsomere 1 almost as long as metatarsomeres 2-4 combined, metatarsomere 5 longer than metatarsomere 2.

Abdomen from visible tergite V slightly narrowed towards both base and apex. Elevated area between two basal lines on first three visible tergites very densely and coarsely punctate. Punctation of all tergites dense and slightly finer than that on elytra. Surface without microsculpture. Setation similar to those on elytra.

Male. Protarsomeres 1-3 strongly dilated, each densely covered with modified pale setae ventrally, protarsomere 4 distinctly narrower than preceding ones, heart-shaped. Aedeagus ( Figs. 25-28 ).

Female. Protarsomeres 1-3 moderately dilated, slightly sub-bilobed, each covered with modified pale setae ventrally, protarsomere 4 scarcely dilated, distinctly narrower than preceding ones.

Differential diagnosis. Philonthus madianus is very similar to P. peripateticus but it differs by the slenderer and longer antennae, wider head and by the different shape of the aedeagus.

Remarks. It is not necessary to designate a lectotype in this case, because evidently Berhauer clearly separated between ‘Typus’ and ‘Cotypus’, which translates to holotype and paratype of the modern concept.

Bionomics. Unknown.

Distribution. Angola, Uganda ( BERNHAUER 1936, HERMAN 2001).

Figures

Figs. 20-28. 20-24 – Philonthus lamtoensis Levasseur, 1967; 25-28 – P. madianus Bernhauer, 1936. 20-22, 25- 27 – aedeagus: 20, 25 ventral view, 21, 26 – dorsal view, 22, 27 – lateral view; 23, 28 – undersite of paramere, with sensory peg setae; 24 – apical portion of sternite VIII, ventral view.