Squatina tergocellatoides (Chen, 1963)

Jonathan H. Walsh & David A. Ebert, 2007, A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squati, Zootaxa 1551, pp. 31-47: 45

publication ID


publication LSID


persistent identifier


treatment provided by


scientific name

Squatina tergocellatoides (Chen, 1963)


Squatina tergocellatoides (Chen, 1963)  ZBK 

Squatina tergocellatoides, Chen 1963  ZBK  : 99, Fig 28, valid, holotype (unique): THUP 00348 (Eschmeyer, 2005). Figure 6.

Common name. Ocellated angel shark

Etymology. Named in reference to the patterns of ocelli on the pectoral fins.

Distribution. Endemic to the WNP including the south China Sea, and waters surrounding northern Taiwan (Compagno et al., 2005a), and northwestern Malaysia (Yano et al., 2005).

Remarks. Additional pictures from Yano et al. (2005) confirm that the coloration and dorsal origin posterior of the pelvic fin tips are diagnostic for this species. There are two other features that are potential characters for S. tergocellatoides  ZBK  present in photographs but require further confirmation. Examination of photographs of fresh S. tergocellatoides  ZBK  specimens suggests that the caudal fin has a unique triangular shape to WNP squatinids in the ventral lobe of the caudal fin. Additionally, Compagno (in press) suggested that the nasal barbels are particularly ornate in S. tergocellatoides  ZBK  , which also is evident in photographs from Yano et al. (2005). Due to a lack of specimens, these characters could not be confirmed in this study.

Attempts to examine or obtain photographs the S. tergocellatoides  ZBK  holotype for examination were unsuccessful; the holotype (THUP 00348) is believed to be lost.


Of the four reported WNP squatinids, Squatina japonica  ZBK  and S. tergocellatoides  ZBK  are distinct from the other two WNP species in that their pelvic fin tips do not reach the origin of the first dorsal fin base. Unique characters distinguishing S. japonica  ZBK  and S. tergocellatoides  ZBK  from one another include a prominent row of midback thorns and unique dorsal coloration (Fig 4) for S. japonica  ZBK  and the distinct paired ocellus pattern (Fig 6), which is exclusive to S. tergocellatoides  ZBK  . Additional distinguishing characters for S. tergocellatoides  ZBK  are the fringing on the nasal barbels and the caudal fin shape, but more photographs or specimens should be examined to confirm these characters.