Panjange kapit Huber

Bernhard A. Huber & Charles Leh Moi Ung, 2016, The Panjange nigrifrons group in Borneo (Araneae: Pholcidae): high diversity in Sarawak, apparent absence in Sabah, European Journal of Taxonomy 184 (184), pp. 1-32 : 16-19

Panjange kapit Huber , sp. nov.

urn:lsid:zoobank.org:act:07838D0A-C949-4316-8B22-6892CC371A3F

Figs 4 , 10–14 , 33–37 , 42–51 , 58–60

Panjange Bor 78: Huber & Nuñeza 2015: 5, 43–44.

Diagnosis

Distinguished from similar species (with pair of straight pointed processes arising from near PME and ventral apophysis on male palpal femur: Pa. bako ; Pa. niah sp. nov.; Pa. pueh sp. nov.; Pa. sedgwicki ; Pa. tahai comb. nov.) by longer palpal segments (especially femur and procursus; Figs 33–34 ), by shapes of procursus and appendix ( Figs 33–34 ), and by longer epigynal scape ( Figs 36–37 ; female of Pa. sedgwicki unknown); from very similar Pa. sedgwicki (male holotype examined) by distal structures of procursus, more slender male palpal tibia, and more distal ventral femur apophysis.

Etymology

Named for the type locality; noun in apposition.

Type material

MALAYSIA-BORNEO: holotype, Ƌ, Sarawak, near Kapit, forest along river (1.937– 1.944° N, 112.904– 112.910° E), 80–120 m a.s.l, 20 Jul. 2014 (B.A. Huber, S.B. Huber), ZFMK ( Ar 14582 ) .

Other material examined

MALAYSIA-BORNEO: Sarawak: 8 ƋƋ, 6 ♀♀, 2 juvs, same data as holotype, ZFMK ( Ar 14583-84 ; 7 ƋƋ, 5 ♀♀) and SMK (1 Ƌ, 1 ♀) ; 1 Ƌ, 3 ♀♀, 1 juv., same data, in absolute ethanol, ZFMK ( Bor 184 ) .

Description

Male (holotype)

MEASUREMENTS. Total body length 5.2, carapace width 1.2. Leg 1: 49.9 (11.1 + 0.5 + 11.6 + 23.8 + 2.9), tibia 2: 7.0, tibia 3: 4.1, tibia 4: 6.3; tibia 1 L/d: 110. Distance PME–PME 520 µm, diameter PME 115 µm, distance PME–ALE ~45 µm; AME absent.

COLOR. Carapace pale ochre yellow to whitish, posterior mark ( Fig. 11 ) lost in ethanol, ocular area and clypeus dark brown (black in life), without black spots in AME area; sternum whitish; legs ochre-orange with dark brown patellae and tibia-metatarsus joints; abdomen ochre-gray, with black marks dorsally, monochromous ventrally.

BODY. Habitus as in Figs 10–11 ; ocular area raised, each triad on long stalk, with pointed straight process arising from near PME ( Fig. 4 ); carapace without median furrow; clypeus unmodified; sternum wider than long (0.75/0.55), unmodified. ALS as in Fig. 44 .

CHELICERAE. Similar to close relatives, with proximal pair of processes and with distinctively bifid distal apophyses ( Figs 35 , 46–47 ); without modified hairs; without stridulatory ridges.

PALPS. As in Figs 33–34 ; coxa unmodified; trochanter with slightly curved pointed retrolatero-ventral apophysis; femur with ventral apophysis; procursus very long, with row of about 18 ventral ridges ( Fig. 49 ) and isolated patch of few prolateral ridges, with distinctive distal processes ( Figs 48–49, 51 ), apparently without process in distal pit ( Fig. 42 ); bulb with strong proximal sclerite, long slender appendix with hooked tip ( Fig. 50 ), and long partly sclerotized embolus with short distal fringes ( Fig. 51 ).

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with> 40 indistinct pseudosegments.

Variation

Tibia 1 in 8 other males: 10.6–11.9 (mean 11.3). Some males with black spots in AME area; abdominal marks variably distinct.

Female

In general similar to male but eye triads on low humps, much closer together (distance PME–PME 220 µm), without pointed processes. Tibia 1 in 6 females: 7.9–8.4 (mean 8.2). Epigynum weakly sclerotized plate with variably sclerotized posterior ‘knob’ on extensible scape ( Figs 36 , 58 ), internal anterior sclerite and complex transversal folds visible through cuticle; internal genitalia as in Figs 37 , 59–60 .

Natural history

Webs were found among vegetation close to the ground, with the apex of the domed sheet connected to the underside of a leaf. Large numbers of Cecidomyiidae (deposited in ZFMK) were seen hanging from the silk lines in most webs. Egg sacs were slightly elongated, and contained about 25– 30 eggs each (n = 2).

Distribution

Known from type locality in Sarawak only ( Fig. 1 ).

Figures

Fig. 1. Known distribution of the Panjange nigrifrons group.

Figs 2 – 7. Male prosomata, oblique frontal views. 2. Panjange pueh Huber, sp. nov. (ZFMK Ar 14576). 3. Pa. niah Huber, sp. nov. (ZFMK Ar 14579). 4. Pa. kapit Huber, sp. nov. (ZFMK Ar 14583). 5. Pa. nigrifrons Deeleman-Reinhold & Deeleman, 1983 (RMNH, Sepaku). 6. Pa. kubah Huber, sp. nov. (ZFMK Ar 14587). 7. Pa. seowi Huber, sp. nov. (ZFMK Ar 14590).

Figs 8 – 15. Live specimens. 8 – 9. Panjange pueh Huber, sp. nov., Ƌ and ♀ from Pueh. 10 – 14. Pa. kapit Huber, sp. nov. from Kapit. 10 – 11. Adult Ƌ, 12. Penultimate instar Ƌ, 13 – 14. ♀ with egg-sac. 15. Pa. niah Huber, sp. nov., ♀ with egg-sac from Niah.

Figs 33 – 34. Panjange kapit Huber, sp. nov. (ZFMK Ar 14583). Left male palp, prolateral and retrolateral views. Abbreviations: a = appendix; b = genital bulb; e = embolus; f = femur; p = procursus; ps = proximal bulbal sclerite; tr = trochanter. Scale line: 1 mm.

Figs 35 – 37. Panjange kapit Huber, sp. nov. (ZFMK Ar 14583 – 84). 35. Male ocular area, clypeus, and chelicerae, oblique frontal view. 36 – 37. Cleared female genitalia, ventral and dorsal views. Scale lines: 0.5 mm.

Figs 38 – 45. — 38 – 41. Panjange niah Huber, sp. nov. (ZFMK Ar 14580). 38. Female prosoma, frontal view. 39 – 40. Comb-hairs on female tarsus 4. 41. Epigynum, ventral view. — 42 – 45. Panjange kapit Huber, sp. nov. (ZFMK Ar 14583). 42. Tip of left procursus, retrolatero-distal view. 43. Base of palpal trichobothrium. 44. Male ALS. 45. Comb-hairs on male tarsus 4. Scale lines: 10 µm (40, 43 – 44); 20 µm (45); 40 µm (39); 60 µm (42); 200 µm (38, 41).

Figs 46 – 51. Panjange kapit Huber, sp. nov. (ZFMK Ar 14583). 46. Male prosoma, frontal view. 47. Right distal male cheliceral apophysis. 48. Right procursus (and bulbal processes), dorsal view. 49. Ridges on left procursus, prolateral view. 50. Tip of left appendix, prolateral view. 51. Tip of left procursus (and embolus), ventral view. Abbreviations: a = appendix; e = embolus; p = procursus. Scale lines: 30 µm (47); 60 µm (50); 100 µm (49, 51); 200 µm (48); 400 µm (46).

Figs 52 – 60. Female genitalia, untreated in ventral view, cleared in ventral and dorsal views. 52 – 54. Panjange pueh Huber, sp. nov. (ZFMK Ar 14577). 55 – 57. Pa. niah Huber, sp. nov. (ZFMK Ar 14580). 58 – 60. Pa. kapit Huber, sp. nov. (ZFMK Ar 14584).