Antithala neugeboreni ( Hoernes & Auinger, 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 12

Antithala neugeboreni ( Hoernes & Auinger, 1880) nov. comb.

Figs 5A 1 –A 2 , B 1 –B 2 , C 1 –C 2

[ Callithea ] Neugeboreni nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra Neugeboreni nov. form.— Hoernes & Auinger 1880: 89, pl. 10, figs 23–24.

Mitra (Costellaria) annae n. sp. — Boettger 1902: 14, nr. 42.

Vexillum (Costellaria) annae (Boettger) — Zilch 1934: 259, pl. 17, fig. 17.

Type material. Lectotype (designated herein): NHMW 1863 /0040/0005, SL: 6.1 mm, MD: 2.7 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 10, figs 24a–c), figs 5A 1 –A 2 . Paralectotypes: NHMW 2020 /0055/0001, SL: 5.8 mm, MD: 2.3 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 10, figs 23a–c), figs 5B 1 –B 2 ; NHMW 2020 /0055/0002, SL: 6.2 mm, MD: 2.1 mm, Lăpugiu de Sus ( Romania), figs 5C 1 –C 2; 5 specimens , NHMW 2020 /0055/0003, Lăpugiu de Sus ( Romania) .

Additional material. SMF 360348 (= SMF XII 2215 a), lectotype of Mitra (Costellaria) annae Boettger, 1902 , illustrated by Zilch (1934, pl. 22, fig. 9), Senckenberg Museum, Frankfurt/Main, Germany, Coşteiu de Sus (Romania).

Revised description. Shell small to medium-sized, broad fusiform with slightly gradate spire and weakly undulating suture. Protoconch largely missing, last protoconch whorl high, smooth. Teleoconch of 5.5 straight-sided to weakly convex whorls. Axial sculpture of spire whorls of prominent axial ribs separated by narrower interspaces, intersected by three weak spiral grooves forming four rows of subquadrate rounded nodes at intersections. Subsutural row of nodes more prominent, separated from following spiral rows by slightly deeper spiral groove. A narrow secondary spiral row of elongate nodes is intercalated in this groove in some specimens. Last whorl strongly constricted with distinct basal concavity and long base, bearing 10–12 spiral rows of nodes plus four spirals on fasciole; spiral sculpture usually weaker than axial ribs; nodes often reduced. Aperture elongate, narrow with distinct, narrow anal sinus, bordered by weak parietal denticle. Columella narrow, sharply delimited, bearing four folds, weakening abapically. Outer lip thickened with up to ten denticles of variable strength within. Siphonal canal moderately long, straight or bent slightly to left, with shallow siphonal notch.

Shell measurements and ratios. SL: 5.8–6.2 mm, MD: 2.3–2.9 mm, AA: 41°, SL/MD: 2.4–2.6, AL/AW: 7.2–7.4, AH/S: 2.1–2.4.

Discussion. Antithala neugeboreni ( Hoernes & Auinger, 1880) can be differentiated from A. sturi ( Hoernes & Auinger, 1880) by its broader outline, gradate spire and the reduced nodes on the last whorl, resulting in dominant axial sculpture. Mitra annae was based by Boettger (1902) on two specimens from Coşteiu de Sus. Boettger (1902) recognized the similarity with A. neugeboreni , but separated M. annae based on the weak spiral sculpture on the last whorl. The reduction of spiral elements is typical in A. neugeboreni and, after having studied the lectotype of M annae , we consider it to be simply a morphotype of A. neugeboreni .

Palaeoenvironment. Unknown; the collections from Lăpugiu de Sus indicate a mixing of coastal-inner neritic and middle neritic elements whereas the assemblages from Coşteiu de Sus suggest middle neritic palaeoenvironments with soft bottom (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 5. A1–A2. Antithala neugeboreni (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW1863/0040/0005. B1–B2. Antithala neugeboreni (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Roma- nia), 2020/0055/0001. C1–C2. Antithala neugeboreni (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 2020/0055/0002. D1–D2. Antithala sturi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. E1–E2. Antithala sturi (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 1863/0015/0155. F1–F2. Antithala sturi (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 2020/0058/0003.