Antithala sturi ( Hoernes & Auinger, 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 12-14

Antithala sturi ( Hoernes & Auinger, 1880) nov. comb.

Figs 4B , 5D 1 –D 2 , E 1 –E 2 , F 1 –F 2

[ Callithea ] Sturi nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra Sturi nov. form.— Hoernes & Auinger 1880: 90 (pars), pl. 10, figs 25–27 [non figs 28–31 = Antithala filipescui nov. sp.].

Mitra (Thala) sturi Hö. Au. — Boettger 1906: 11.

Thala Sturi R. Hoern. i Auing— Friedberg 1911: 31, pl. 1, fig. 24.

Thala Sturi R. Hoern. i Auing—Friedberg 1928: 580, pl. 37, figs 30–31.

Th [ala]. sturi (R. Hörn, et Au.)— Sieber 1958a: 153.

Thala sturi (Hoern. et Auing.) — Kókay 1966: 62, pl. 9, fig. 6.

Vexillum (Uromitra) sturi ( Hoernes & Auinger, 1880) — Atanacković 1985: 161, pl. 36, figs 5–6.

Thala sturi ( Hoernes & Auinger, 1880) — Bałuk 1997: 39, pl. 11, figs 4–6.

Type material. Lectotype (designated herein): NHMW 2020 /0058/0001, SL: 5.3 mm, MD: 1.9 mm, Lăpugiu de Sus ( Romania), figs 4B, 5D 1 –D 2 . Paralectotypes: NHMW 1863 /0015/0155, SL: 4.6 mm, MD: 1.9 mm, Lăpugiu de Sus ( Romania) , illustrated in Hoernes & Auinger (1880, pl. 10, figs 27a–c), figs 5E 1 –E 2; NHMW 2020 /0058/0003, SL: 5.9 mm, MD: 2.2 mm, Lăpugiu de Sus ( Romania) , illustrated in Hoernes & Auinger (1880, pl. 10, figs 26a–c), figs 5F 1 –F 2; NHMW 2020 /0056/0003, Coşteiu de Sus ( Romania) ; NHMW 2020 /0058/0002, Lăpugiu de Sus ( Romania) , illustrated in Hoernes & Auinger (1880, pl. 10, figs 25a–c). 15 specimens, NHMW 2020 /0058/0004, Lăpugiu de Sus ( Romania) .

Revised description. Shell small, moderately slender, fusiform, with conical to faintly cyrtoconoid spire. Protoconch (fig. 4B) high conical of three weakly convex, smooth whorls (height: 530 μm, diameter: 450 μm). Teleoconch of 4.5 straight-sided whorls, with periphery at abapical suture, separated by deeply incised suture. First teleoconch whorl with blunt, weakly opisthocline axial ribs separated by deep, narrow interspaces. Axial ribs intersected by two weaker spiral grooves resulting in three spiral rows of prominent, subquadrate nodes on spire whorls. Fourth spiral row of nodes intercalated on third and fourth teleoconch whorls along abapical suture, but largely covered by succeeding whorl. Axial and spiral interspaces deeply cut, resulting in regular, prominent sculpture of square nodes. Last whorl moderately convex, moderately constricted base, with nine spiral rows of nodes axially arranged in weakly sigmoidal ribs. About six prominent spiral cords with strongly reduced nodes on fasciole. Aperture elongate, narrow with distinct anal sinus narrowed by broad, low parietal denticle. Columella narrow, sharply delimited, bearing three prominent folds, weakening abapically, plus weaker fourth fold bordering siphonal canal. Outer lip strongly thickened, with five to six stout denticles within. Siphonal canal moderately long, bent slightly to left, with shallow siphonal notch.

Shell measurements and ratios. SL: 4.6–5.9 mm, MD: 1.9–2.2 mm; AA: 40°, SL/MD: 2.7–2.9, AL/AW: 5.7–7.9, AH/S: 2.2–2.3.

Discussion. Antihala sturi ( Hoernes & Auinger, 1880) is characterised by its regularly beaded sculpture and displays little variability in shape. Some specimens develop a slightly wider spiral groove below the first adapical spiral cord (e.g. Bałuk 1997, pl. 11, figs 4–6). We are not aware of any similar species from the European Neogene except for the Paratethyan species discussed herein.

Palaeoenvironment. Unknown; the collections from Lăpugiu de Sus indicate a mixing of coastal-inner neritic and middle neritic elements.

Distribution in Central Paratethys. Badenian (middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1997); Ukrainian Fore-Carpathian Basin: Zborów (Zboriv) ( Ukraine); ( Friedberg 1911,); Pannonian Basin: Herend ( Hungary) ( Kókay 1966); southern Pannonian Basin: Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); Făget Basin: Lăpugiu de Sus (own data), Coşteiu de Sus ( Romania) ( Boettger 1906).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes & Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes & Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037.

FIGURE 5. A1–A2. Antithala neugeboreni (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW1863/0040/0005. B1–B2. Antithala neugeboreni (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Roma- nia), 2020/0055/0001. C1–C2. Antithala neugeboreni (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 2020/0055/0002. D1–D2. Antithala sturi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. E1–E2. Antithala sturi (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 1863/0015/0155. F1–F2. Antithala sturi (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 2020/0058/0003.