Bellardithala lapugyensis ( Hoernes & Auinger, 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 25-26

Bellardithala lapugyensis ( Hoernes & Auinger, 1880) nov. comb.

Figs 9A 1 –A 2 , B 1 –B, C 1 –C 2

[ Callithea ] Lapugyensis nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra Lapugyensis nov. form.— Hoernes & Auinger 1880 (pars): 89, pl. 10, figs 21a–c [non pl. 10., figs 22a–22c = Bellardithala fedosovi nov. sp.].

Mitra (Thala) lapugyensis Hö. Au. — Boettger 1906: 11.

? Turricula Lapugyensis R. H. i Auing.—Friedberg 1928: 579, pl. 37, fig. 28.

Thala lapugyensis ( Hoernes & Auinger, 1880) — Bałuk 1997: 38, pl. 11, fig. 1.

Type material. Lectotype (designated herein): NHMW 1854 /0035/0101, SL: 5.4 mm, MD: 2.0 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 10, figs 21a–c), figs 9A 1 –A 2 . Paralectotypes: NHMW 2020 /0053/0001, SL: 5.1 mm, MD: 2.0 mm, Lăpugiu de Sus ( Romania), figs 9B 1 –B 2 ; NHMW 2020 /0053/0002, SL: 5.4 mm, MD: 2.1 mm, Lăpugiu de Sus ( Romania), figs 9C 1 –C 2; 2 specimens , NHMW 2020 /0053/0003, 2 specimens , NHMW 2020 /0053/0002, all Lăpugiu de Sus ( Romania) .

Revised description. Shell small, moderately broad fusiform with weakly cyrtoconoid spire. Protoconch high conical with 2.7 regularly convex whorls (height: 500 μm, diameter: 510 μm). Transition to teleoconch marked by onset of axial sculpture. Teleoconch of 4.5 weakly convex whorls, with periphery close above abapical suture, separated by deeply incised suture. Axial sculpture of about 15 prominent axial ribs on first teleoconch whorl, separated by slightly narrower interspaces. Spiral sculpture consisting of indistinct spiral grooves separating four spiral rows of blurred nodules. Cancellate sculpture on later whorls, with axial ribs and interspaces of similar width, crossed by four broad, regularly spaced spiral cords on second teleoconch whorl and five on subsequent whorl. Spiral cords form low subquadrate nodules at intersection with axial ribs, and well defined grooves in interspaces. Abapically, spiral interspaces reduced to rectangular pits. Last whorl weakly convex, with weakly constricted base, bearing about 13 spiral cords; 7 spirals on weak fasciole. Aperture narrow, elongate, with indistinct anal sinus. Columellar callus very narrow, indistinct. Columella with four columellar folds, weakening abapically. Outer lip only slightly thickened, bearing eight small denticles within. Siphonal canal moderately long, moderately wide, straight or bent slightly left, with wide, shallow siphonal notch.

Shell measurements and ratios. SL: 5.1–5.4 mm, MD: 2.0– 2.1 mm, AA: 40–43°, SL/MD: 2.5–2.7, AL/AW: 4.1–4.8, AH/S: 2.3–2.4.

Discussion. Hoernes & Auinger (1880) based the description of Mitra lapugyensis on specimens from Lăpugiu de Sus ( Romania) and from Steinebrunn ( Austria). In our opinion they are not conspecific, and as first revisers we designate the specimen illustrated by Hoernes & Auinger (1880, pl. 10, fig. 21) from Lăpugiu de Sus as lectotype. The specimen from Steinebrunn illustrated by Hoernes & Auinger (1880, pl. 10, fig. 22) is described herein as Bellardithala fedosovi nov. sp. Bellardithala lapugyensis differs from congeneric Paratethyan species in its cancellate sculpture and small size. The specimen described by Friedberg (1928) from the Badenian of Ukraine differs from typical M. lapugyensis in its more slender outline and the narrower spiral cords. It is doubtful that it is conspecific with B. lapugyensis .

Palaeoenvironment. Unknown; the collections from Lăpugiu de Sus indicate a mixing of coastal-inner neritic and middle neritic elements.

Distribution in Central Paratethys. Badenian (middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1997),? Ukrainian Fore-Carpathian Basin: Zborów (Zboriv) ( Ukraine) (Friedberg 1928); Făget Basin: Lăpugiu de Sus ( Romania) ( Hoernes & Auinger 1880).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 9 A1–A2. Bellardithala lapugyensis (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0101. B1–B. Bellardithala lapugyensis (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 2020/0053/0001. C1–C2. Bellardithala lapugyensis (Hoernes & Auinger, 1880), paralectotype, Lăpugiu de Sus (Roma- nia), 2020/0053/0002. D1–D2. Bellardithala partschi (Hörnes, 1852), lectotype, Steinebrunn (Austria), NHMW 1846/0037/0113. E1–E2. Bellardithala partschi (Hörnes, 1852), paralectotype, Steinebrunn (Austria), NHMW 2020/0060/0004. F1–F2. Bellardithala partschi (Hörnes, 1852), paralectotype, Steinebrunn (Austria), NHMW 1860/0001/0110.