Ebenomitra pseudopyramidella ( Boettger, 1906 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 31-33

Ebenomitra pseudopyramidella ( Boettger, 1906) nov. comb.

Figs 4F , 10D 1 –D 2 , E 1 –E 2 , F 1 –F 2 , 11B 1 –B 2

V.[oluta] laevis n.— Eichwald 1829: 297, pl. 5, fig. 14 [non Donovan, 1804].

Mitra laevis m.— Eichwald 1830: 221.

Mitra laevis Eichw. — Dubois de Montpereux 1831: 25, pl. 1, figs 2–3.

Mitra turgidula m. [sic]—Pusch 1837: 118, pl. 11, fig. 4.

Mitra laevis Dubois — Pusch 1837: 119.

[ Mitra incognita ] var. β laevis m.—Pusch 1837: 119, pl.11, fig. 5.

Mitra laevis — Eichwald 1840: 12.

Mitra laevis — Eichwald 1852: pl. 8, fig. 17.

Mitra laevis m.— Eichwald 1853: 203.

Mitra (Volutomitra) ebenus Lamk. — Hoernes & Auinger 1880: 82, pl. 7, fig. 13 [non Lamarck, 1811].

* Mitra (Uromitra) ebenus Lmk. var. pseudopyramidella n. nom.— Boettger 1906: 9 [nov. nom. pro Mitra ebenus Hoernes & Auinger, 1880 pl. 7, fig. 13].

Turricula ebenus var. striata Eichw. — Friedberg 1911: 22, pl. 1, figs 14–15.

Turricula ebenus var. laevis . Eichw.— Friedberg 1911: 23, pl. 1, fig. 16.

Mitra laevis Eichw. — Hilber 1882: 2, pl. 1, fig. 1.

Type material. Holotype: NHMW 2020 /0071/0001, SL: 16.9 mm, MD: 6.3 mm, Ternopil ( Ukraine), illustrated in Hoernes & Auinger (1880, pl. 7, fig. 13), figs 10E 1 –E 2.

Additional material. NHMW 2020 /0071/0002, SL: 17.0 mm, MD: 6.5 mm, Ternopil ( Ukraine) , figs 10F 1 –F 2; NHMW 2020 /0071/0003, SL: 15.8 mm, MD: 6.1 mm, Ternopil ( Ukraine) , figs 10D 1 –D 2; NHMW 2020 /0071/0004, SL: 16.1 mm, MD: 5.6 mm, Ternopil ( Ukraine) , figs 11B 1 –B 2; NHMW 1857 /0014/0009, SL: 16.5 mm, MD: 6.1 mm, Steinebrunn ( Austria) , fig. 4F; NHMW 2020 /0071/0005, 8 specimens, Ternopil ( Ukraine) ; NHMW 1859 /0040/0006, 1 specimen. Zaleśce ( Ukraine) ; NHMW 1857 /0014/0009, 51 specimens, Steinebrunn ( Austria) .

Revised description. Shell moderately large, solid, glossy, moderately broad fusiform, with high, conical to weakly cyrtoconoid spire. Protoconch (fig. 4F) paucispiral, with low nucleus and about two weakly convex whorls (height: 540 μm, diameter: 530 μm) (note that transition to teleoconch is abraded in available specimen overemphasizing the weak convexity of the last protoconch whorl). Teleoconch of about eight weakly convex to subcylindrical whorls, with periphery at or slightly above abapical suture. Suture narrowly impressed, slightly undulating. Axial sculpture on early teleoconch whorls consisting of low, convex axial ribs, separated by narrower interspaces. Axial sculpture fading out within fourth to fifth whorl, rarely persisting on penultimate whorl. Last whorl moderately contracting. Aperture wide, ovoid, with indistinct anal sinus. Columellar callus narrow, sharply delimited. Columella with three oblique folds; adapical two folds most prominent. Outer lip with 12–15 lirae placed distinctly behind peristome; lirae more prominent at their adapertural termination, fading out inside. Outer lip weakly thickened at lirae, thinning towards peristome. Siphonal canal moderately wide, short, straight or weakly bent to left. Spiral sculpture absent, except for few poorly defined spiral cords on weak fasciole; most prominent spiral cord extending from adapical columellar fold. Colour pattern frequently preserved, consisting of orange-brown background with narrow white band slightly below mid-whorl on spire whorls and above periphery on last whorl.

Shell measurements and ratios. SL: 12.6–17.0 mm, MD: 4.5–6.5 mm; AA: 38–42°, AA: 45–52°, SL/MD: 2.5–2.7, AL/AW: 3.2–3.5, AH/S: 3.1–3.4.

Discussion. Eichwald (1829, 1853) described Voluta laevis from Biłka, Poczajów Stary, Żukowice, Tarnoruda and Zaleśce in Ukraine and Hilber (1882) added material from Hołubica ( Ukraine). Boettger (1906) introduced Mitra pseudopyramidella as new name for a specimen described by Hoernes & Auinger (1880 pl. 7, fig. 13) from Ternopil, situated close to the other Ukrainian localities. The shells from Ternopil are clearly conspecific with ‘ Mitra laevis ’ specimens from Hołubica, Żukowice and Zaleśce in the NHMW and GBA collections. Therefore, we treat both taxa as synonyms. Voluta laevis is preoccupied by Donovan (1804) and therefore, Mitra pseudopyramidella Boettger, 1906 is the next available name. Pusch (1837) discussed this species as Mitra turgidula , referring to Voluta turgidula Brocchi, 1814 from the Pliocene of Italy. Thus, although he added “m” (for mihi) when describing the species, he did not intend to establish it as new species.

Ebenomitra pseudopyramidella differs from Pusia avellanella ( Boettger, 1906) in its larger size, glossy shell, and absence of a fourth columellar fold. Ebenomitra sublaevis Landau, Ceulemans & Van Dingenen, 2019 , from the Tortonian of France, is highly reminiscent of E. pseudopyramidella , but differs in its smaller size and stout outline (see Landau et al., 2019, pl. 52, figs 1–4).

Distribution in Central Paratethys. Badenian (middle Miocene): Voronyaky Hills: Hołubica (Holubytsia) ( Ukraine) ( Friedberg 1911); Ukrainian Fore-Carpathian Basin: Dryszczów (Nadrichne), Tarnoruda (Tarnoruda), Zaleśce (Salisze) ( Ukraine), Zborów (Zboriv), Żukowce (Zhukivtsi) ( Ukraine) ( Friedberg 1911); Vienna Basin: Steinebrunn ( Austria).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes & Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes & Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037.

FIGURE 10 A1–A2. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1870/0033/0034. B1– B2. Ebenomitra leucozona (Andrzejowski, 1830), Gainfarn (Austria), NHMW 1846/0037/0110. C1–C2. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1868/0001/0401. D1–D2. Ebenomitra pseudopyramidella (Boettger, 1906), Ternopil (Ukraine), NHMW 2020/0071/0003. E1–E2. Ebenomitra pseudopyramidella (Boettger, 1906), holotype, Ternopil (Ukraine), NHMW 2020/0071/0001. F1–F2. Ebenomitra pseudopyramidella (Boettger, 1906), Ternopil (Ukraine), NHMW 2020/0071/0002. G1–G2. Ebenomitra januszkiewiczi (Friedberg, 1928), lectotype, Zboriv (Zborów) (Ukraine), PAN A-I-50/35. H1–H2. Ebenomitra januszkiewiczi (Friedberg, 1928), paralectotype, Zboriv (Zborów) (Ukraine), ZNG PAN A-I-50/36.1-36.6. I1–I2. Ebenomitra januszkiewiczi (Friedberg, 1928), paralectotype, Zboriv (Zborów) (Ukraine), ZNG PAN A-I-50/36.1-36.6. J. Ebenomitra januszkiewiczi (Friedberg, 1928), paralectotype, Zboriv (Zborów) (Ukraine). ZNG PAN A-I-50/36.1-36.6.

FIGURE 11 A1–A2. Ebenomitra leucozona (Andrzejowski, 1830), Steinebrunn (Austria), NHMW 1857/0014/0008. B1–B2. Ebenomitra pseudopyramidella (Boettger, 1906), Ternopil (Ukraine), NHMW 2020/0071/0004.