Pusia? falsitranssylvanica, 1840

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 39

Pusia? falsitranssylvanica nov. nom.

Figs 13G

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Mitra (Costellaria) recticosta var. transsylvanica n.— Boettger, 1902: 13 [non Hoernes & Auinger 1880].

Mitra (Costellaria) pseudorecticosta var. transsylvanica Bttgr. — Boettger, 1906: 10 [non Hoernes & Auinger 1880].

Vexillum (Costellaria) pseudorecticosta var. transsylvanica (Boettger) — Zilch 1934: 258, pl. 17, fig. 15 [non Hoernes & Auinger 1880].

Type material. Holotype: SMF 360346 (= XII.2212 a), Senckenberg Museum, Frankfurt/Main, Germany, SL: 7.3 mm, MD: 2.2 mm, Coşteiu de Sus ( Romania), illustrated in Zilch (1934, pl. 17, fig. 15), figs 13G 1 –G 2.

Type locality. Coşteiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, Badenian.

Etymology. A combination of falsus (Latin for wrong) and transsylvanica .

Diagnosis. Pusia species of medium-size, moderately slender fusiform profile, with high spire, convex spire whorls, convex last whorl, and prominent, widely spaced opisthocline axial ribs.

Description. Medium sized, slender fusiform, glossy with high spire. Protoconch largely broken, last protoconch whorl convex, smooth (diameter: 0.5 mm). Teleoconch of six convex whorls, with periphery slightly below mid-whorl, separated by incised suture. Axial sculpture of prominent, opisthocline, convex ribs, separated by slightly wider, concave interspaces (14 axial ribs on penultimate whorl); ribs weakening towards adapical suture, prominent at abapical suture, undulating suture. Last whorl convex, constricted at base, bearing sigmoidal axial ribs, fading over base. Four prominent spiral cords on fasciole. Aperture wide; columellar callus indistinct; columella with three prominent folds. Outer lip not preserved; no lirae inside aperture. Siphonal canal moderately short, moderately wide, straight, with very shallow siphonal notch.

Shell measurements and ratios. SL: 7.3 mm, MD: 2.2 mm; AA: 31°, AA: 45–52°, SL/MD: 2.9, AL/AW: 3.4, AH/S: 2.8.

Discussion. This species was treated by Boettger (1902, 1906) as variety transsylvanica of Mitra recticosta sensu Hoernes & Auinger, 1880 [= Pusia pseudorecticosta ( Boettger, 1906) ]. Mitra (Costellaria) recticosta transsylvanica Boettger, 1902 , however, is a primary homonym of Mitra (Cylindra) transsylvanica Hoernes & Auinger 1880 , and we therefore introduce falsitranssylvanica as replacement name.

Mitra (Costellaria) recticosta transsylvanica Boettger, 1902 (= Pusia falsitranssylvanica ), was treated as senior synonym of Mitra pseudorecticosta Boettger, 1906 by Landau et al. (2013). Pusia falsitranssylvanica differs distinctly from P. pseudorecticosta in its convex whorls (instead of subcylindrical) and the more prominent, wider spaced, opisthocline axial ribs. Moreover, the occurrence at Coşteiu de Sus suggests a preference for middle neritic soft bottom environments, whereas P. pseudorecticosta is typically found in silty-sandy deposits of coastal environments. The placement in Pusia is tentative.

Palaeoenvironment. The assemblages from Coşteiu de Sus suggest middle neritic palaeoenvironments with soft bottom (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1906).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 13 A1–A2. Pusia avellanella (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360343. B1–B2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0002. C1–C2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. D1–D2. Pusia avellanella (Boettger, 1906), Coşteiu de Sus (Romania), SMF 360344. E1–E2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0004. F1–F2. Pusia avellanella (Boettger, 1906), Steinebrunn (Austria), NHMW 1847/0037/0048. G1–G2. Pusia falsitranssylvanica nov. nom., holotype, Coşteiu de Sus (Roma- nia), SMF 360346. H1–H2. Pusia confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. I1–I2. Pusia moravica (Hoernes & Auinger, 1880), lectotype, Rudice (Czech Republic), NHMW 1862/0029/0006. J. Pusia moravica (Hoernes & Auinger, 1880), paralectotype, Rudice (Czech Republic), NHMW 2020/0097/0001.